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Shirley, W. D.; Leung, Y.-M. (1970). Medusae of the Central Arctic. In Taxonomic Guides to Arctic Zooplankton. II. University of Southern California Los Angeles, Department of Biological Sciences, Los Angeles. 3: 3-18.
Shirley, W. D.; Leung, Y.-M. (1970). Medusae of the Central Arctic. In Taxonomic Guides to Arctic Zooplankton. II. University of Southern California Los Angeles, Department of Biological Sciences, Los Angeles. 3: 3-18.
