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Harl J., Haring E., Asami T., Sittenthaler M., Sattmann H. & Páll-Gergely B. (2017). Molecular systematics of the land snail family Orculidae reveal paraphyly and deep splits within the clade Orthurethra (Gastropoda: Pulmonata). Zoological Journal of the Linnean Society. 181(4): 778-794.