According to article 68.2.1. of the ICZN, "
The expressions "gen. n., sp. n.", "new genus and species", or an equivalent, applied before 1931 to only one of two or more new nominal species originally included in a new nominal genus or subgenus, are deemed to be an original designation if no other type species was explicitly designated." This way,
Lycastoides alticola Johnson, 1903 is the type of the genus
Lycastoides by original designation.