Mantelatto, F.L.; Robles, R.; Schubart, C.D.; Felder, D.L. (2009). Molecular phylogeny of the genus
Cronius Stimpson, 1860, with reassignment of
C. tumidulus and several American species of
Portunus to the genus
Achelous De Haan, 1833 (Brachyura: Portunidae).
In: Crustacean Issues 18: Decapod Crustacean Phylogenetics, Martin, J.W., Crandall, K.A. & Felder, D.L. (eds). CRC Press, England, pp. 567–579.