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WoRMS taxon details

Varunidae H. Milne Edwards, 1853

439140  (urn:lsid:marinespecies.org:taxname:439140)

  1. Biota
  2. Animalia (Kingdom)
  3. Arthropoda (Phylum)
  4. Crustacea (Subphylum)
  5. Multicrustacea (Superclass)
  6. Malacostraca (Class)
  7. Eumalacostraca (Subclass)
  8. Eucarida (Superorder)
  9. Decapoda (Order)
  10. Pleocyemata (Suborder)
  11. Brachyura (Infraorder)
  12. Eubrachyura (Section)
  13. Thoracotremata (Subsection)
  14. Grapsoidea (Superfamily)
  15. Varunidae (Family)
accepted
Family

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  1. Subfamily Asthenognathinae Stimpson, 1858
    1. Genus Asthenognathus Stimpson, 1858
      1. Species Asthenognathus alleronensis Pasini, Garassino & De Angeli, 2017 †
      2. Species Asthenognathus atlanticus Monod, 1933
      3. Species Asthenognathus australensis Feldmann, Schweitzer, Casadío & Griffin, 2011 †
      4. Species Asthenognathus cornishorum Schweitzer & Feldmann, 1999 †
      5. Species Asthenognathus fernandezi Ferratges, Zamora & Aurell, 2022 †
      6. Species Asthenognathus globosa (Karasawa, 1990) †
      7. Species Asthenognathus hexagonum Rathbun, 1909
      8. Species Asthenognathus inaequipes Stimpson, 1858
      9. Species Asthenognathus laverdensis De Angeli & Garassino, 2006 †
      10. Species Asthenognathus microspinus Casadío, De Angeli, Feldmann, Garassino, Hetler, Parras & Schweitzer, 2004 †
      11. Species Asthenognathus rakosensis Müller, 2006 †
      12. Species Asthenognathus sakumotoi Karasawa, 2018 †
      13. Species Asthenognathus urretae Schweitzer & Feldmann, 2001 †
      14. Species Asthenognathus gallardoi Serène & Soh, 1976 accepted as Gopkittisak gallardoi (Serène & Soh, 1976) (unaccepted > superseded combination)
    2. Genus Globihexapus Schweitzer & Feldmann, 2001 †
      1. Species Globihexapus kosekii Kato, 2005 †
      2. Species Globihexapus paxillus Schweitzer & Feldmann, 2001 †
    3. Tribe Gustavini Števčić, 2011 accepted as Aphanodactylidae Ahyong & Ng, 2009
  2. Subfamily Cyclograpsinae H. Milne Edwards, 1853
    1. Genus Austrohelice Sakai, Türkay & Yang, 2006
      1. Species Austrohelice crassa (Dana, 1851)
      2. Species Austrohelice manneringi Feldmann, Schweitzer, Maxwell & Kelley, 2008 †
    2. Genus Chasmagnathus De Haan, 1833
      1. Species Chasmagnathus convexus (De Haan, 1835)
      2. Species Chasmagnathus convexus Haswell, 1881 accepted as Helograpsus haswellianus (Whitelegge, 1890) (unaccepted > junior homonym)
      3. Species Chasmagnathus georgei Clark, 1987 † accepted as Parahelice georgei (Clark, 1987) (unaccepted > superseded combination)
      4. Species Chasmagnathus granulatus Dana, 1851 accepted as Neohelice granulata (Dana, 1851)
      5. Species Chasmagnathus Haswellianus Whitelegge, 1890 accepted as Helograpsus haswellianus (Whitelegge, 1890) (unaccepted > superseded combination)
      6. Species Chasmagnathus laevis Dana, 1851 accepted as Paragrapsus laevis (Dana, 1851) (unaccepted > superseded combination)
      7. Species Chasmagnathus subquadratus Dana, 1851 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > superseded combination)
    3. Genus Cyclograpsus H. Milne Edwards, 1837
      1. Species Cyclograpsus audouinii H. Milne Edwards, 1837
      2. Species Cyclograpsus barbatus (MacLeay, 1838)
      3. Species Cyclograpsus beccarii Nobili, 1899
      4. Species Cyclograpsus cinereus Dana, 1851
      5. Species Cyclograpsus directus Karasawa, 1989 †
      6. Species Cyclograpsus escondidensis Rathbun in Glassell, 1933
      7. Species Cyclograpsus eydouxi H. Milne Edwards, 1853
      8. Species Cyclograpsus granulatus Dana, 1851
      9. Species Cyclograpsus granulosus H. Milne Edwards, 1853
      10. Species Cyclograpsus henshawi Rathbun, 1902
      11. Species Cyclograpsus incisus Shen, 1940
      12. Species Cyclograpsus insularum Campbell & Griffin, 1966
      13. Species Cyclograpsus integer H. Milne Edwards, 1837
      14. Species Cyclograpsus intermedius Ortmann, 1894
      15. Species Cyclograpsus lavauxi H. Milne Edwards, 1853
      16. Species Cyclograpsus longipes Stimpson, 1858
      17. Species Cyclograpsus lucidus Dai in Dai, Yang, Song & Chen, 1986
      18. Species Cyclograpsus lwangi Hu & Tao, 1999 †
      19. Species Cyclograpsus minutus Hombron & Jacquinot, 1846
      20. Species Cyclograpsus pumilio Hangai & Fukui in Hangai, Kitaura, Wada & Fukui, 2009
      21. Species Cyclograpsus punctatus H. Milne Edwards, 1837
      22. Species Cyclograpsus rectangularis Karasawa, 1989 †
      23. Species Cyclograpsus sanctaecrucis Griffin, 1968
      24. Species Cyclograpsus unidens Nobili, 1905
      25. Species Cyclograpsus audini H. Milne Edwards, 1837 accepted as Cyclograpsus audouinii H. Milne Edwards, 1837 (unaccepted > unjustified emendation)
      26. Species Cyclograpsus crenulatus H. Milne Edwards, 1837 accepted as Hemigrapsus crenulatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      27. Species Cyclograpsus gaimardii H. Milne Edwards, 1837 accepted as Paragrapsus gaimardii (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      28. Species Cyclograpsus gibbus Hombron & Jacquinot, 1846 accepted as Hemigrapsus gibbus (Hombron & Jacquinot, 1846) (unaccepted > superseded combination)
      29. Species Cyclograpsus gnatherion Kinahan, 1857 accepted as Pinnotherelia laevigata H. Milne Edwards & Lucas, 1843 (unaccepted > junior subjective synonym)
      30. Species Cyclograpsus laevis Hess, 1865 accepted as Cyclograpsus audouinii H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
      31. Species Cyclograpsus latreillii H. Milne Edwards, 1837 accepted as Pseudohelice latreillii (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      32. Species Cyclograpsus lophopus Nobili, 1906 accepted as Cyclorma lophopum (Nobili, 1906)
      33. Species Cyclograpsus marmoratus White, 1847 (unaccepted > nomen nudum)
      34. Species Cyclograpsus occidentalis A. Milne-Edwards, 1878 accepted as Cyclograpsus integer H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
      35. Species Cyclograpsus octodentatus H. Milne Edwards, 1837 accepted as Leptograpsodes octodentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      36. Species Cyclograpsus parvulus De Man, 1896 accepted as Cyclograpsus integer H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
        1. Variety Cyclograpsus parvulus var. unidens Nobili, 1905 accepted as Cyclograpsus unidens Nobili, 1905 (unaccepted > superseded combination)
      37. Species Cyclograpsus quadridentatus H. Milne Edwards, 1837 accepted as Paragrapsus quadridentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      38. Species Cyclograpsus reynaudi H. Milne Edwards, 1853 accepted as Cyclograpsus punctatus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
      39. Species Cyclograpsus sexdentatus H. Milne Edwards, 1837 accepted as Hemigrapsus sexdentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      40. Species Cyclograpsus whitei H. Milne Edwards, 1853 accepted as Cyclograpsus lavauxi H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
      41. Species Cyclograpsus tasmanicus Hombron & Jacquinot, 1846 (uncertain > taxon inquirendum)
    4. Genus Helicana Sakai & Yatsuzuka, 1980
      1. Species Helicana doerjesi Sakai, Türkay & Yang, 2006
      2. Species Helicana japonica (Sakai & Yatsuzuka, 1980)
      3. Species Helicana wuana (Rathbun, 1931)
    5. Genus Helice De Haan, 1833
      1. Species Helice epicure Ng, Naruse & Shih, 2018
      2. Species Helice formosensis Rathbun, 1931
      3. Species Helice latimera Parisi, 1918
      4. Species Helice tientsinensis Rathbun, 1931
      5. Species Helice tridens (De Haan, 1835)
        1. Subspecies Helice tridens formosensis Rathbun, 1931 accepted as Helice formosensis Rathbun, 1931 (unaccepted > superseded combination)
        2. Subspecies Helice tridens pingi Rathbun, 1931 accepted as Helice latimera Parisi, 1918 (unaccepted > junior subjective synonym)
        3. Subspecies Helice tridens sheni Sakai, 1939 accepted as Helicana wuana (Rathbun, 1931) (unaccepted > junior subjective synonym)
        4. Subspecies Helice tridens tientsinensis Rathbun, 1931 accepted as Helice tientsinensis Rathbun, 1931 (unaccepted > superseded combination)
        5. Variety Helice tridens var. latimera Parisi, 1918 accepted as Helice latimera Parisi, 1918 (unaccepted > superseded combination)
        6. Subspecies Helice tridens wuana Rathbun, 1931 accepted as Helice wuana Rathbun, 1931 (unaccepted > superseded combination)
      6. Subgenus Helice (Helicana) Sakai & Yatsuzuka, 1980 accepted as Helicana Sakai & Yatsuzuka, 1980 (status change)
        1. Species Helice (Helicana) japonica Sakai & Yatsuzuka, 1980 accepted as Helicana japonica (Sakai & Yatsuzuka, 1980) (unaccepted > superseded combination)
      7. Species Helice crassa Dana, 1851 accepted as Austrohelice crassa (Dana, 1851) (unaccepted > superseded combination)
      8. Species Helice dentipes Heller, 1865 accepted as Metaplax dentipes (Heller, 1865) (unaccepted > superseded combination)
      9. Species Helice gaudichaudi H. Milne Edwards, 1853 accepted as Neohelice granulata (Dana, 1851) (unaccepted > junior subjective synonym)
      10. Species Helice japonica Sakai & Yatsuzuka, 1980 accepted as Helicana japonica (Sakai & Yatsuzuka, 1980) (unaccepted > superseded combination)
      11. Species Helice latreillei (H. Milne Edwards, 1837) accepted as Helice tridens (De Haan, 1835) (unaccepted > junior subjective synonym, unjustified emendation)
      12. Species Helice latreilli (H. Milne Edwards, 1837) accepted as Helice tridens (De Haan, 1835) (unaccepted > junior subjective synonym, unjustified emendation)
      13. Species Helice leachi Hess, 1865 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > misspelling - incorrect subsequent spelling)
      14. Species Helice Leachii Hess, 1865 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > junior subjective synonym)
      15. Species Helice lucasi H. Milne Edwards, 1853 accepted as Austrohelice crassa (Dana, 1851) (unaccepted > junior subjective synonym)
      16. Species Helice pilimana A. Milne-Edwards, 1873 accepted as Parahelice pilimana (A. Milne-Edwards, 1873) (unaccepted > superseded combination)
      17. Species Helice spinicarpa H. Milne Edwards, 1853 accepted as Chasmagnathus convexus (De Haan, 1835) (unaccepted > junior subjective synonym)
      18. Species Helice wuana Rathbun, 1931 accepted as Helicana wuana (Rathbun, 1931) (unaccepted > superseded combination)
    6. Genus Helograpsus Campbell & Griffin, 1966
      1. Species Helograpsus haswellianus (Whitelegge, 1890)
    7. Genus Metaplax H. Milne Edwards, 1852
      1. Species Metaplax crenulata (Gerstaecker, 1856)
      2. Species Metaplax dentipes (Heller, 1865)
      3. Species Metaplax distincta H. Milne Edwards, 1852
      4. Species Metaplax elegans De Man, 1888
      5. Species Metaplax gocongensis Davie & V.X. Nguyen, 2003
      6. Species Metaplax indica H. Milne Edwards, 1852
      7. Species Metaplax intermedia De Man, 1888
      8. Species Metaplax longipes Stimpson, 1858
      9. Species Metaplax occidentalis Pretzmann, 1971
      10. Species Metaplax sheni Gordon, 1930
      11. Species Metaplax tredecim Tweedie, 1950
      12. Species Metaplax crassipes De Man, 1892 accepted as Metaplax elegans De Man, 1888 (unaccepted > junior subjective synonym)
      13. Species Metaplax distinctus H. Milne Edwards, 1852 accepted as Metaplax distincta H. Milne Edwards, 1852 (unaccepted > incorrect grammatical agreement of specific epithet)
      14. Species Metaplax hirsutimana Grant & McCulloch, 1906 accepted as Australoplax tridentata (A. Milne-Edwards, 1873) (unaccepted > junior subjective synonym)
      15. Species Metaplax hirtipes Heller, 1862 accepted as Hemiplax hirtipes (Hombron & Jacquinot, 1846) (unaccepted > junior subjective synonym)
      16. Species Metaplax indicus H. Milne Edwards, 1852 accepted as Metaplax indica H. Milne Edwards, 1852 (unaccepted > incorrect grammatical agreement of specific epithet)
        1. Subspecies Metaplax indicus occidentalis Pretzmann, 1971 accepted as Metaplax occidentalis Pretzmann, 1971 (unaccepted > superseded combination)
      17. Species Metaplax intermedius De Man, 1888 accepted as Metaplax intermedia De Man, 1888 (unaccepted > superseded combination)
      18. Species Metaplax takahashii Sakai, 1939 accepted as Metaplax longipes Stimpson, 1858 (unaccepted > junior subjective synonym)
    8. Genus Miosesarma Karasawa, 1989 †
      1. Species Miosesarma japonica Karasawa, 1989 †
      2. Species Miosesarma nagurense Kato, 1996 †
    9. Genus Neohelice Sakai, Türkay & Yang, 2006
      1. Species Neohelice granulata (Dana, 1851)
    10. Genus Paragrapsus H. Milne Edwards, 1853
      1. Species Paragrapsus gaimardii (H. Milne Edwards, 1837)
      2. Species Paragrapsus laevis (Dana, 1851)
      3. Species Paragrapsus quadridentatus (H. Milne Edwards, 1837)
      4. Species Paragrapsus urvillei H. Milne Edwards, 1853
      5. Species Paragrapsus verreauxi H. Milne Edwards, 1853 accepted as Paragrapsus laevis (Dana, 1851) (unaccepted > junior subjective synonym)
    11. Genus Parahelice Sakai, Türkay & Yang, 2006
      1. Species Parahelice balssi (Sakai, Türkay & Yang, 2006)
      2. Species Parahelice daviei (Sakai, Türkay & Yang, 2006)
      3. Species Parahelice georgei (Clark, 1987)
      4. Species Parahelice ngankeeae Hsu & Shih, 2024
      5. Species Parahelice pilimana (A. Milne-Edwards, 1873)
      6. Species Parahelice pilosa (Sakai, Türkay & Yang, 2006)
    12. Genus Pseudohelice Sakai, Türkay & Yang, 2006
      1. Species Pseudohelice annamalai Prema, Hsu, Shih & Ravichandran, 2022
      2. Species Pseudohelice latreillii (H. Milne Edwards, 1837)
      3. Species Pseudohelice subquadrata (Dana, 1851)
      4. Subgenus Pseudohelice (Parahelice) Sakai, Türkay & Yang, 2006 accepted as Parahelice Sakai, Türkay & Yang, 2006
        1. Species Pseudohelice (Parahelice) balssi Sakai, Türkay & Yang, 2006 accepted as Parahelice balssi (Sakai, Türkay & Yang, 2006) (unaccepted > superseded combination)
        2. Species Pseudohelice (Parahelice) daviei Sakai, Türkay & Yang, 2006 accepted as Parahelice daviei (Sakai, Türkay & Yang, 2006) (unaccepted > superseded combination)
        3. Species Pseudohelice (Parahelice) pilosa Sakai, Türkay & Yang, 2006 accepted as Parahelice pilosa (Sakai, Türkay & Yang, 2006)
    13. Genus Gnathochasmus MacLeay, 1838 accepted as Cyclograpsus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
      1. Species Gnathochasmus barbatus MacLeay, 1838 accepted as Cyclograpsus barbatus (MacLeay, 1838) (unaccepted > superseded combination)
    14. Genus Rhaconotus Gerstaecker, 1856 accepted as Metaplax H. Milne Edwards, 1852 (name pre-occupied by Rhaconotus Ruthe, 1854 [Hymenoptera])
      1. Species Rhaconotus crenulatus Gerstaecker, 1856 accepted as Metaplax crenulata (Gerstaecker, 1856) (unaccepted > superseded combination)
  3. Subfamily Gaeticinae Davie & Ng, 2007
    1. Genus Acmaeopleura Stimpson, 1858
      1. Species Acmaeopleura parvula Stimpson, 1858
      2. Species Acmaeopleura rotunda Rathbun, 1909
      3. Species Acmaeopleura balssi Shen, 1932 accepted as Sestrostoma balssi (Shen, 1932) (unaccepted > superseded combination)
      4. Species Acmaeopleura depressa Sakai, 1965 accepted as Sestrostoma depressum (Sakai, 1965) (unaccepted > superseded combination)
      5. Species Acmaeopleura hichiro Karasawa & Tanaka, 2005 † accepted as Sestrostoma hichiro (Karasawa & Tanaka, 2005) † (unaccepted > superseded combination)
      6. Species Acmaeopleura toriumii Takeda, 1974 accepted as Sestrostoma toriumii (Takeda, 1974) (unaccepted > superseded combination)
    2. Genus Brankocleistostoma Števčić, 2011
      1. Species Brankocleistostoma fossulum (Barnard, 1955)
    3. Genus Gaetice Gistel, 1848
      1. Species Gaetice depressus (De Haan, 1833)
      2. Species Gaetice ungulatus Sakai, 1939
    4. Genus Gopkittisak Naruse & Clark, 2009
      1. Species Gopkittisak angustum Komai, 2011
      2. Species Gopkittisak gallardoi (Serène & Soh, 1976)
      3. Species Gopkittisak ngankeeae Lasley, Anker & Naruse, 2024
    5. Genus Paranotonyx Nobili, 1906
      1. Species Paranotonyx curtipes Nobili, 1906
    6. Genus Proexotelson Naruse, 2015
      1. Species Proexotelson tokoroi Naruse, 2015
    7. Genus Pseudopinnixa Ortmann, 1894
      1. Species Pseudopinnixa carinata Ortmann, 1894
    8. Genus Sestrostoma Davie & Ng, 2007
      1. Species Sestrostoma balssi (Shen, 1932)
      2. Species Sestrostoma depressum (Sakai, 1965)
      3. Species Sestrostoma hichiro (Karasawa & Tanaka, 2005) †
      4. Species Sestrostoma toriumii (Takeda, 1974)
    9. Genus Goetice Gistel, 1848 accepted as Gaetice Gistel, 1848 (ICZN Article 33.2.3.1 used to suppress this original spelling in favor of Gaetice (see Ng et al., 2008) )
      1. Species Goetice americanus Rathbun, 1923 accepted as Pseudogaetice americanus (Rathbun, 1923) (unaccepted > superseded combination)
    10. Tribe Gopkittisakini Števčić, 2011 accepted as Gaeticinae Davie & Ng, 2007
    11. Genus Plalynotus De Haan, 1835 accepted as Platynotus De Haan, 1835 (unaccepted > misspelling - incorrect subsequent spelling)
    12. Genus Platygrapsus Stimpson, 1858 accepted as Gaetice Gistel, 1848 (unnecessary replacement name for Grapsus (Platynotus) De Haan, 1833)
      1. Species Platygrapsus convexiusculus Stimpson, 1858 accepted as Gaetice depressus (De Haan, 1833) (unaccepted > junior subjective synonym)
      2. Species Platygrapsus depressus (De Haan, 1835) accepted as Gaetice depressus (De Haan, 1833) (unaccepted > superseded combination)
    13. Genus Platynotus De Haan, 1835 accepted as Gaetice Gistel, 1848 (unaccepted > junior homonym)
      1. Species Platynotus depressus (De Haan, 1833) accepted as Gaetice depressus (De Haan, 1833) (unaccepted > superseded combination)
  4. Genus Hemiplax Heller, 1865
    1. Species Hemiplax hirtipes (Hombron & Jacquinot, 1846)
    2. Species Hemiplax major Glaessner, 1960 †
    3. Species Hemiplax boteltobagoe Sakai, 1939 accepted as Macrophthalmus (Paramareotis) boteltobagoe (Sakai, 1939) (unaccepted > superseded combination)
    4. Species Hemiplax hirtipes (Heller, 1862) accepted as Hemiplax hirtipes (Hombron & Jacquinot, 1846) (unaccepted > junior subjective synonym, also junior secondary homonym)
  5. Subfamily Pinnotherelinae Alcock, 1900
    1. Genus Pinnotherelia H. Milne Edwards & Lucas, 1843
      1. Species Pinnotherelia laevigata H. Milne Edwards & Lucas, 1843
    2. Genus Viapinnixa Schweitzer & Feldmann, 2001 †
      1. Species Viapinnixa alvarezi Vega, Cosma, Coutiño, Feldmann, Nyborg, Schweitzer & Waugh, 2001 †
      2. Species Viapinnixa mexiaensis Armstrong, Nyborg, Bishop, Ossó-Morales & Vega, 2009 †
      3. Species Viapinnixa nodosa (Collins & Rasmussen, 1992) †
      4. Species Viapinnixa perrilliatae Vega, Nyborg, Fraaye & Espinosa, 2007 †
  6. Subfamily Thalassograpsinae Davie & Ng, 2007
    1. Genus Thalassograpsus Tweedie, 1950
      1. Species Thalassograpsus harpax (Hilgendorf, 1892)
  7. Genus Tritodynamia Ortmann, 1894
    1. Species Tritodynamia bengalensis Trivedi, Mitra & Ng, 2021
    2. Species Tritodynamia bidentata Yang & Tang, 2005
    3. Species Tritodynamia dilatata Yang & Sun, 1996
    4. Species Tritodynamia fujianensis Chen, 1979
    5. Species Tritodynamia hainanensis Dai in Dai, Feng, Song & Chen, 1980
    6. Species Tritodynamia horvathi Nobili, 1905
    7. Species Tritodynamia intermedia Shen, 1935
    8. Species Tritodynamia japonica Ortmann, 1894
    9. Species Tritodynamia longipropoda Dai in Dai, Feng, Song & Chen, 1980
    10. Species Tritodynamia miocaenica Müller, 2006 †
    11. Species Tritodynamia nontjii Rahayu & Ng, 2021
    12. Species Tritodynamia rathbuni Shen, 1932
    13. Species Tritodynamia serratipes Anker & Ng, 2014
    14. Species Tritodynamia wangi Ng & Ho, 2023
    15. Species Tritodynamia yeoi Naruse & Ng, 2010
    16. Species Tritodynamia dilatatum Yang & Sun, 1996 accepted as Tritodynamia dilatata Yang & Sun, 1996 (unaccepted > incorrect grammatical agreement of specific epithet)
    17. Species Tritodynamia fani Shen, 1932 accepted as Tritodynamia horvathi Nobili, 1905 (unaccepted > junior subjective synonym)
    18. Species Tritodynamia globosa Karasawa, 1990 † accepted as Asthenognathus globosa (Karasawa, 1990) † (unaccepted > superseded combination)
    19. Species Tritodynamia longipropodum Dai in Dai, Feng, Song & Chen, 1980 accepted as Tritodynamia longipropoda Dai in Dai, Feng, Song & Chen, 1980 (unaccepted > incorrect grammatical agreement of specific epithet)
  8. Subfamily Varuninae H. Milne Edwards, 1853
    1. Genus Brachynotus De Haan, 1833
      1. Species Brachynotus atlanticus Forest, 1957
      2. Species Brachynotus corallinus Beschin, Busulini, De Angeli & Tessier, 2007 †
      3. Species Brachynotus februarius Müller, 1974 †
      4. Species Brachynotus foresti Zariquiey Álvarez, 1968
      5. Species Brachynotus gemmellari (Rizza, 1839)
      6. Species Brachynotus multituberculatus Beschin, Busulini & Tessier in Beschin, Busulini, Tessier & Zorzin, 2019 †
      7. Species Brachynotus oligocenicus De Angeli, Garassino & Ceccon, 2010 †
      8. Species Brachynotus sexdentatus (Risso, 1827)
      9. Subgenus Brachynotus (Heterograpsus) Lucas, 1846 accepted as Brachynotus De Haan, 1833 (unaccepted > junior subjective synonym)
        1. Species Brachynotus (Heterograpsus) jouyi Rathbun, 1894 accepted as Tetragrapsus jouyi (Rathbun, 1894) (unaccepted > superseded combination)
      10. Species Brachynotus brevidigitatus Yokoya, 1928 accepted as Hemigrapsus penicillatus (De Haan, 1835) (unaccepted > junior subjective synonym)
      11. Species Brachynotus edwardsii Hilgendorf, 1882 accepted as Hemigrapsus sexdentatus (H. Milne Edwards, 1837) (unaccepted > junior subjective synonym)
      12. Species Brachynotus gemmellaroi (Rizza, 1839) accepted as Brachynotus gemmellari (Rizza, 1839) (unaccepted > misspelling - incorrect subsequent spelling)
      13. Species Brachynotus harpax Hilgendorf, 1892 accepted as Thalassograpsus harpax (Hilgendorf, 1892) (unaccepted > superseded combination)
      14. Species Brachynotus longitarsis (Miers, 1879) accepted as Hemigrapsus longitarsis (Miers, 1879) (unaccepted > superseded combination)
      15. Species Brachynotus lucasi (H. Milne Edwards, 1853) accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > junior subjective synonym)
      16. Species Brachynotus marmoratus (Olivi, 1792) accepted as Pachygrapsus marmoratus (Fabricius, 1787) (unaccepted > superseded combination)
      17. Species Brachynotus penicillatus (De Haan, 1835) accepted as Hemigrapsus penicillatus (De Haan, 1835) (unaccepted > superseded combination)
      18. Species Brachynotus spinosus (H. Milne Edwards, 1853) accepted as Parvonotus spinosus (H. Milne Edwards, 1853) (unaccepted > superseded combination)
    2. Genus Chhapgarus Ng, Trivedi & Bhat, 2022
      1. Species Chhapgarus intermedius (Chhapgar, 1957)
    3. Genus Cyrtograpsus Dana, 1851
      1. Species Cyrtograpsus affinis (Dana, 1851)
      2. Species Cyrtograpsus altimanus Rathbun, 1914
      3. Species Cyrtograpsus angulatus Dana, 1851
      4. Species Cyrtograpsus cirripes Smith, 1869
    4. Genus Eoacmaeopleura De Angeli & Bellin, 2022 †
      1. Species Eoacmaeopleura arzignanensis De Angeli & Bellin, 2022 †
    5. Genus Eriocheir De Haan, 1835
      1. Species Eriocheir hepuensis Dai, 1991
      2. Species Eriocheir japonica (De Haan, 1835)
      3. Species Eriocheir ogasawaraensis Komai in Komai, Yamasaki, Kobayashi, Yamamoto & Watanabe, 2006
      4. Species Eriocheir sinensis H. Milne Edwards, 1853
        1. Forma Eriocheir sinensis f. acutifrons Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
        2. Forma Eriocheir sinensis f. rostrata Panning, 1933 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior objective synonym)
        3. Forma Eriocheir sinensis f. rotundifrons Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
        4. Forma Eriocheir sinensis f. trilobata Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
      5. Species Eriocheir formosa Chan, Hung & Yu, 1995 accepted as Platyeriocheir formosa (Chan, Hung & Yu, 1995) (unaccepted > superseded combination)
      6. Species Eriocheir formosa Nakagawa, 1915 accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > nomen nudum)
      7. Species Eriocheir japonicus (De Haan, 1835) accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > misspelling - incorrect subsequent spelling)
        1. Subspecies Eriocheir japonicus hepuensis Dai, 1991 accepted as Eriocheir hepuensis Dai, 1991 (unaccepted > superseded combination)
      8. Species Eriocheir leptognathus Rathbun, 1913 accepted as Neoeriocheir leptognathus (Rathbun, 1913) (unaccepted > superseded combination)
      9. Species Eriocheir misakiensis Rathbun, 1919 accepted as Hemigrapsus longitarsis (Miers, 1879) (unaccepted > junior subjective synonym)
      10. Species Eriocheir rectus Stimpson, 1858 accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > junior subjective synonym)
    6. Genus Grapsodius Holmes, 1900
      1. Species Grapsodius eximius Holmes, 1900
    7. Genus Hemigrapsus Dana, 1851
      1. Species Hemigrapsus crassimanus Dana, 1851
      2. Species Hemigrapsus crenulatus (H. Milne Edwards, 1837)
      3. Species Hemigrapsus estellinensis Creel, 1964
      4. Species Hemigrapsus gibbus (Hombron & Jacquinot, 1846)
      5. Species Hemigrapsus longitarsis (Miers, 1879)
      6. Species Hemigrapsus nudus (Dana, 1851)
      7. Species Hemigrapsus oregonensis (Dana, 1851)
      8. Species Hemigrapsus pallipes (H. Milne Edwards, 1837)
      9. Species Hemigrapsus penicillatus (De Haan, 1835)
      10. Species Hemigrapsus sanguineus (De Haan, 1835)
      11. Species Hemigrapsus sexdentatus (H. Milne Edwards, 1837)
      12. Species Hemigrapsus sinensis Rathbun, 1931
      13. Species Hemigrapsus takanoi Asakura & Watanabe, 2005
      14. Species Hemigrapsus affinis Dana, 1851 accepted as Cyrtograpsus affinis (Dana, 1851) (unaccepted > superseded combination)
      15. Species Hemigrapsus lobulatus Manuel, Gonzales & Basmayor, 1991 accepted as Noarograpsus lobulatus (Manuel, Gonzales & Basmayor, 1991) (unaccepted > superseded combination)
      16. Species Hemigrapsus tanakoi Asakura & Watanabe, 2005 accepted as Hemigrapsus takanoi Asakura & Watanabe, 2005 (unaccepted > misspelling - incorrect subsequent spelling)
    8. Genus Neoeriocheir Sakai, 1983
      1. Species Neoeriocheir leptognathus (Rathbun, 1913)
    9. Genus Noarograpsus Ng, Manuel & Ng, 2006
      1. Species Noarograpsus lobulatus (Manuel, Gonzales & Basmayor, 1991)
    10. Genus Orcovita Ng & Tomascik, 1994
      1. Species Orcovita angulata Ng, Guinot & Iliffe, 1996
      2. Species Orcovita fictilia Ng, Guinot & Iliffe, 1996
      3. Species Orcovita gracilipes Ng, Guinot & Iliffe, 1996
      4. Species Orcovita hicksi Davie & Ng, 2012
      5. Species Orcovita holthuisi Ng & Ng, 2009
      6. Species Orcovita mcneiceae Ng & Ng, 2002
      7. Species Orcovita miruku Naruse & Tamura, 2006
      8. Species Orcovita mollitia Ng, Guinot & Iliffe, 1996
      9. Species Orcovita orchardorum Davie & Ng, 2012
      10. Species Orcovita saltatrix Ng & Tomascik, 1994
      11. Species Orcovita tabiacoud Stasolla & Innocenti, 2014
      12. Species Orcovita varungan Ng & Naruse, 2024
    11. Genus Otognathon Ng & Števčić, 1993
      1. Species Otognathon komodoense (Serène, 1971)
      2. Species Otognathon uru Ng, Komai & Ng, 2009
    12. Genus Parapyxidognathus Ward, 1941
      1. Species Parapyxidognathus deianira (De Man, 1888)
      2. Species Parapyxidognathus fluviatilis (Alcock, 1900)
      3. Species Parapyxidognathus ongia Hsu & Shih, 2024
      4. Species Parapyxidognathus tshiansue Hsu & Shih, 2024
    13. Genus Parvonotus Davie & Ng, 2024
      1. Species Parvonotus meridionalis Davie & Ng, 2024
      2. Species Parvonotus samoensis Davie & Ng, 2024
      3. Species Parvonotus spinosus (H. Milne Edwards, 1853)
    14. Genus Platyeriocheir Ng, Guo & Ng, 1999
      1. Species Platyeriocheir formosa (Chan, Hung & Yu, 1995)
      2. Species Platyeriocheir guangdonga Sakai, 2013 accepted as Platyeriocheir formosa (Chan, Hung & Yu, 1995) (unaccepted > junior subjective synonym)
    15. Genus Pseudogaetice Davie & Ng, 2007
      1. Species Pseudogaetice americanus (Rathbun, 1923)
    16. Genus Pseudograpsus H. Milne Edwards, 1837
      1. Species Pseudograpsus albus Stimpson, 1858
      2. Species Pseudograpsus crassus A. Milne-Edwards, 1868
      3. Species Pseudograpsus elongatus (A. Milne-Edwards, 1873)
      4. Species Pseudograpsus setosus (Fabricius, 1798)
      5. Species Pseudograpsus barbatus H. Milne Edwards, 1853 accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
      6. Species Pseudograpsus dentatus Miers, 1880 accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
      7. Species Pseudograpsus erythraeus Kossman, 1877 accepted as Pseudograpsus elongatus (A. Milne-Edwards, 1873) (unaccepted > junior subjective synonym)
      8. Species Pseudograpsus intermedius Chhapgar, 1957 accepted as Chhapgarus intermedius (Chhapgar, 1957) (unaccepted > superseded combination)
      9. Species Pseudograpsus laniger Tesch, 1918 accepted as Scutumara laniger (Tesch, 1918) (unaccepted > superseded combination)
      10. Species Pseudograpsus miyakei Nakamura & Kurata, 1972 accepted as Scutumara miyakei (Nakamura & Takeda, 1972) (unaccepted > superseded combination)
      11. Species Pseudograpsus nudus Dana, 1851 accepted as Hemigrapsus nudus (Dana, 1851) (unaccepted > superseded combination)
      12. Species Pseudograpsus oregonensis Dana, 1851 accepted as Hemigrapsus oregonensis (Dana, 1851) (unaccepted > superseded combination)
      13. Species Pseudograpsus pallipes H. Milne Edwards, 1837 accepted as Hemigrapsus pallipes (H. Milne Edwards, 1837) (unaccepted > superseded combination)
      14. Species Pseudograpsus penicilliger (Latreille, 1817) accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
    17. Genus Ptychognathus Stimpson, 1858
      1. Species Ptychognathus affinis De Man, 1895
      2. Species Ptychognathus altimanus (Rathbun, 1914)
      3. Species Ptychognathus amikee Hsu & Shih, 2024
      4. Species Ptychognathus andamanensis Pretzmann, 1984
      5. Species Ptychognathus barbatus (A. Milne-Edwards, 1873)
      6. Species Ptychognathus capillidigitatus Takeda, 1984
      7. Species Ptychognathus crassimanus Finnegan, 1931
      8. Species Ptychognathus demani Roux, 1917
      9. Species Ptychognathus dentatus De Man, 1892
      10. Species Ptychognathus easteranus Rathbun, 1907
      11. Species Ptychognathus glaber Stimpson, 1858
        1. Subspecies Ptychognathus glaber andamanesis Pretzmann, 1984 accepted as Ptychognathus andamanensis Pretzmann, 1984 (unaccepted > superseded combination)
      12. Species Ptychognathus guijulugani Rathbun, 1914
      13. Species Ptychognathus hachijoensis Sakai, 1955
      14. Species Ptychognathus insolitus Osawa & Ng, 2006
      15. Species Ptychognathus intermedius (De Man, 1879)
      16. Species Ptychognathus ishii Sakai, 1939
      17. Species Ptychognathus johannae Rathbun, 1913
      18. Species Ptychognathus lipkei Ng, 2010
      19. Species Ptychognathus makii Hsu & Shih, 2020
      20. Species Ptychognathus ngankeeae de Mazancourt, Mazel, Marquet, Poupin & Keith, 2024
      21. Species Ptychognathus onyx Alcock, 1900
      22. Species Ptychognathus pilipes (A Milne-Edwards, 1868)
      23. Species Ptychognathus pilosus De Man, 1892
      24. Species Ptychognathus polleni De Man, 1895
      25. Species Ptychognathus pusillus Heller, 1865
      26. Species Ptychognathus riedelii (A Milne-Edwards, 1868)
        1. Variety Ptychognathus riedelii var. pilosa De Man, 1892 accepted as Ptychognathus pilosus De Man, 1892 (unaccepted > superseded combination)
      27. Species Ptychognathus sakaii Hsu, Shih & Li, 2022
      28. Species Ptychognathus similis de Mazancourt, Mazel, Marquet, Poupin & Keith, 2024
      29. Species Ptychognathus spinicarpus Ortmann, 1894
      30. Species Ptychognathus stimpsoni Hsu & Shih, 2020
      31. Species Ptychognathus takahasii Sakai, 1939
      32. Species Ptychognathus andamanica Alcock, 1900 accepted as Ptychognathus riedelii (A Milne-Edwards, 1868) (unaccepted > junior subjective synonym)
      33. Species Ptychognathus easterana Rathbun, 1907 accepted as Ptychognathus easteranus Rathbun, 1907 (unaccepted > superseded combination)
      34. Species Ptychognathus takahashii Sakai, 1939 accepted as Ptychognathus takahasii Sakai, 1939 (unaccepted > misspelling - incorrect subsequent spelling)
    18. Genus Pyxidognathus A. Milne-Edwards, 1879
      1. Species Pyxidognathus granulosus A. Milne-Edwards, 1879
      2. Species Pyxidognathus subglobosus Tesch, 1918
      3. Species Pyxidognathus deianira De Man, 1888 accepted as Parapyxidognathus deianira (De Man, 1888) (unaccepted > superseded combination)
      4. Species Pyxidognathus deldeni (De Man, 1879) accepted as Pyxidognathus granulosus A. Milne-Edwards, 1879
      5. Species Pyxidognathus fluviatilis Alcock, 1900 accepted as Parapyxidognathus fluviatilis (Alcock, 1900) (unaccepted > superseded combination)
    19. Genus Sakakurapus Karasawa, 2018 †
      1. Species Sakakurapus kogisorum Karasawa, 2018 †
    20. Genus Scutumara Ng & Nakasone, 1993
      1. Species Scutumara enodis Ng & Nakasone, 1993
      2. Species Scutumara laniger (Tesch, 1918)
      3. Species Scutumara miyakei (Nakamura & Takeda, 1972)
    21. Genus Tetragrapsus Rathbun, 1918
      1. Species Tetragrapsus jouyi (Rathbun, 1894)
    22. Genus Utica White, 1847
      1. Species Utica barbimana A. Milne-Edwards, 1873
      2. Species Utica borneensis De Man, 1895
      3. Species Utica crassimana Haswell, 1881
      4. Species Utica glabra A. Milne-Edwards, 1873
      5. Species Utica gracilipes White, 1847
      6. Species Utica haswelli Wintle, 1887 †
      7. Species Utica nausithoe De Man, 1895
      8. Species Utica setosipes Haswell, 1881
      9. Species Utica yarraensis Wintle, 1887 †
      10. Species Utica sinensis Parisi, 1918 accepted as Neoeriocheir leptognathus (Rathbun, 1913) (unaccepted > junior subjective synonym)
    23. Genus Varuna H. Milne Edwards in Bory de Saint Vincent, 1830
      1. Species Varuna litterata (Fabricius, 1798)
      2. Species Varuna yui Hwang & Takeda, 1986
      3. Species Varuna altimana Rathbun, 1914 accepted as Ptychognathus altimanus (Rathbun, 1914) (unaccepted > superseded combination)
      4. Species Varuna angustifrons Karasawa, 1993 † accepted as Carinocarcinoides angustifrons (Karasawa, 1993) † (unaccepted > superseded combination)
      5. Species Varuna atlantica Melliss, 1875 accepted as Planes major (MacLeay, 1838) (unaccepted > junior subjective synonym)
      6. Species Varuna tomentosa Pfeffer, 1889 accepted as Varuna litterata (Fabricius, 1798) (unaccepted > junior subjective synonym)
    24. Genus Coelochirus Nauck, 1880 accepted as Ptychognathus Stimpson, 1858 (unaccepted > junior subjective synonym)
      1. Species Coelochirus crinipes Nauck, 1880 accepted as Ptychognathus pilipes (A Milne-Edwards, 1868) (unaccepted > junior subjective synonym)
    25. Genus Cryptograpsus Dana, 1851 accepted as Cyrtograpsus Dana, 1851 (unaccepted > misspelling - incorrect subsequent spelling)
      1. Species Cryptograpsus cirripes Smith, 1869 accepted as Cyrtograpsus cirripes Smith, 1869 (unaccepted > superseded combination)
    26. Genus Eriochirus H. Milne Edwards, 1853 accepted as Eriocheir De Haan, 1835 (unaccepted > unjustified emendation)
      1. Species Eriochirus japonicus (De Haan, 1835) accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > superseded combination)
      2. Species Eriochirus sinensis H. Milne Edwards, 1853 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > misspelling - incorrect original spelling)
    27. Tribe Euchirograpsini Števčić, 2005 accepted as Plagusiidae Dana, 1851
    28. Genus Gnathograpsus A. Milne-Edwards, 1868 accepted as Ptychognathus Stimpson, 1858 (unaccepted > junior subjective synonym)
      1. Species Gnathograpsus barbatus A. Milne-Edwards, 1873 accepted as Ptychognathus barbatus (A. Milne-Edwards, 1873) (unaccepted > superseded combination)
      2. Species Gnathograpsus intermedius De Man, 1879 accepted as Ptychognathus intermedius (De Man, 1879) (unaccepted > superseded combination)
      3. Species Gnathograpsus pilipes A. Milne-Edwards, 1868 accepted as Ptychognathus pilipes (A Milne-Edwards, 1868) (unaccepted > superseded combination)
      4. Species Gnathograpsus riedelii A. Milne-Edwards, 1868 accepted as Ptychognathus riedelii (A Milne-Edwards, 1868) (unaccepted > superseded combination)
    29. Genus Hypsilograpsus De Man, 1879 accepted as Pyxidognathus A. Milne-Edwards, 1879 (unaccepted > junior subjective synonym)
      1. Species Hypsilograpsus deldeni De Man, 1879 accepted as Pyxidognathus granulosus A. Milne-Edwards, 1879 (unaccepted > junior subjective synonym)
    30. Tribe Otognathini Števčić, 2011 accepted as Varuninae H. Milne Edwards, 1853
    31. Genus Pachystomum Nauck, 1880 accepted as Pseudograpsus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
      1. Species Pachystomum philippinense Nauck, 1880 accepted as Pseudograpsus albus Stimpson, 1858 (unaccepted > junior subjective synonym)
    32. Genus Paraeriocheir Sakai, 2013 accepted as Eriocheir De Haan, 1835 (unaccepted > junior subjective synonym)
    33. Genus Shurebus Verany, 1846 accepted as Brachynotus De Haan, 1833 (unaccepted > junior subjective synonym)
      1. Species Shurebus genoensis White, 1847 accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > nomen nudum)
      2. Species Shurebus genuensis Verany, 1846 accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > junior subjective synonym)
marine, brackish, fresh, terrestrial
Milne Edwards, H. (1853). Mémoire sur la famille des ocypodiens. <em>Annales des Sciences Naturelles, 3rd series.</em> 20: 163-228, pls. 6-11. [details]  OpenAccess publication 
DecaNet eds. (2024). DecaNet. Varunidae H. Milne Edwards, 1853. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=439140 on 2024-07-16
Date
action
by
2009-12-15 09:56:25Z
created
Davie, Peter

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


original description Milne Edwards, H. (1853). Mémoire sur la famille des ocypodiens. <em>Annales des Sciences Naturelles, 3rd series.</em> 20: 163-228, pls. 6-11. [details]  OpenAccess publication 

context source (PeRMS) Moscoso, V. (2012). Catálogo de crustáceos decápodos y estomatópodos del Perú. <em>Boletín Instituto del Mar del Perú.</em> 27(1-2): 8-207., available online at https://repositorio.imarpe.gob.pe/handle/20.500.12958/2190 [details]   
 
 Present  Inaccurate  Introduced: alien  Containing type locality 
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LanguageName 
Japanese モクズガニ科  [details]
To Biological Information System for Marine Life (BISMaL) 
To European Nucleotide Archive, ENA (Varunidae) 
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