WoRMS source details
Korshunova, T.; Fletcher, K.; Martynov, A. (2025). The endless forms are the most differentiated — how taxonomic pseudo-optimization masked natural diversity and evolution: the nudibranch case. Zoological Journal of the Linnean Society. 204(4): zlaf057: 1-93.
514054
10.1093/zoolinnean/zlaf057 [view]
Korshunova, T.; Fletcher, K.; Martynov, A.
2025
The endless forms are the most differentiated — how taxonomic pseudo-optimization masked natural diversity and evolution: the nudibranch case
Zoological Journal of the Linnean Society
204(4): zlaf057: 1-93
Publication
Stated date: 18 August 2025
Available for editors 
[request]
[request]
In this study, a global-encompassing, fine-scale differentiated taxonomy is consistently presented from the high level of a major nudibranch suborder, Aeolidacea, and then consequentially to lower levels of superfamilies, families, genera, and species to conjoin the underlying evolutionary pathways of aeolidacean nudibranchs and systematic representation to a maximal degree. The suborder Aeolidacea is reinstated and the superfamily system of Aeolidacea is reformed; 10 aeolidacean superfamilies are recognized encompassing 29 families in an all-family Synopsis, and the superfamily Embletonioidea is included in addition. Two new families, Chudidae fam. nov. and Hantazuidae fam. nov., are described, which show unique morphological and molecular patterns. Seven new species are described within various superfamilies, and five species are reinstated. The genus-level taxonomy of the family Coryphellidae is revisited, 11 genera are recognized, including nine reinstated and two new genera. We show that recent synonymization of all valid Coryphellidae genera into just a single genus was based on an erroneous phylogenetic analysis, the careless treatment of morphological characters, and established preconceptions. Uniting fine-scale morphological and molecular data, order level nudibranch taxonomy is refined, order Nudibranchia is restricted, and order Doridida is reinstated. Within the order Nudibranchia, five suborders are recognized: Arminacea, restricted, reinstated; Tritoniacea, reinstated; Dendronotacea, restricted, reinstated; Janolacea, reinstated; and Aeolidacea, revised, reinstated.
Date
action
by
Abronicidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Aeolidina (status source)
Amphorina Quatrefages, 1844 (status source)
Amphorina amazighi (Tamsouri, Carmona, Moukrim & Cervera, 2015) (new combination reference)
Amphorina andra Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Amphorina farrani (Alder & Hancock, 1844) (additional source)
Amphorina linensis (García-Gómez, Cervera & F. J. Garcia, 1990) (additional source)
Amphorina pallida (Alder & Hancock, 1842) (additional source)
Amphorina viriola Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Annulorhina K. P. Rao, 1968 (status source)
Annulorhina mandapamensis K. P. Rao, 1968 (status source)
Apataidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Apataoidea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Fionoidea J. E. Gray, 1857 (basis of record)
Arminina (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Borealea nobilis (A. E. Verrill, 1880) accepted as Coryphella nobilis A. E. Verrill, 1880 (status source)
Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Calmidae Iredale & O'Donoghue, 1923 accepted as Flabellinidae Bergh, 1889 (additional source)
Capellinia Trinchese, 1873 (status source)
Capellinia vittata (Alder & Hancock, 1842) (new combination reference)
Catriona aurantia (Alder & Hancock, 1842) (status source)
Chudidae Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo humanistica Korshunova, Fletcher & Martynov, 2025 (original description)
Chudoidea Korshunova, Fletcher & Martynov, 2025 (original description)
Corrupta Korshunova, Fletcher & Martynov, 2025 accepted as Coryphella J. E. Gray, 1850 (original description)
Coryphella alexanderi Ekimova, 2022 (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 (status source)
Coryphella borealis Odhner, 1922 (status source)
Coryphella browni Picton, 1980 (status source)
Coryphella californica Bergh, 1904 (status source)
Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 (status source)
Coryphella falklandica Eliot, 1907 (status source)
Coryphella fisheri MacFarland, 1966 accepted as Orienthella piunca (Er. Marcus, 1961) accepted as Coryphella piunca Er. Marcus, 1961 (source of synonymy)
Coryphella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) (additional source)
Coryphella lineata (Lovén, 1846) (status source)
Coryphella longicaudata O'Donoghue, 1922 (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella nobilis A. E. Verrill, 1880 (additional source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella trilineata O'Donoghue, 1921 (additional source)
Coryphella trophina (Bergh, 1890) (additional source)
Coryphella verta Ev. Marcus, 1970 (additional source)
Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025 (original description)
Cuthona Alder & Hancock, 1855 (additional source)
Cuthonellidae M. C. Miller, 1977 (additional source)
Cuthonidae Odhner, 1934 (additional source)
Dendronotina (status source)
Diaphoreolis flavovulta (MacFarland, 1966) (status source)
Diaphoreolis lagunae (O'Donoghue, 1926) (status source)
Diaphoreolis stipata (Alder & Hancock, 1843) (status source)
Diaphoreolis zvezda Korshunova, Fletcher, Bakken & Martynov, 2023 (status source)
Doridida (status source)
Dunga Eliot, 1902 (status source)
Embletonioidea Pruvot-Fol, 1954 (basis of record)
Eubranchidae Odhner, 1934 (additional source)
Eubranchopsis virginalis Baba, 1949 (status source)
Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchulus olivaceus (O'Donoghue, 1922) (status source)
Eubranchulus rupium (Møller, 1842) (status source)
Eubranchus Forbes, 1838 (additional source)
Eubranchus novik Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024 accepted as Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchus rubropunctatus Edmunds, 1969 accepted as Produnga rubropunctata (Edmunds, 1969) (status source)
Favorinidae Bergh, 1889 accepted as Facelinidae Bergh, 1889 (additional source)
Fionoidea J. E. Gray, 1857 (basis of record)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Fjordia browni (Picton, 1980) accepted as Coryphella browni Picton, 1980 (status source)
Fjordia capensis (Thiele, 1925) accepted as Coryphella capensis Thiele, 1925 (status source)
Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Fjordia insolita (García-Gómez & Cervera, 1989) accepted as Coryphella insolita (García-Gómez & Cervera, 1989) (status source)
Fjordia lineata (Lovén, 1846) accepted as Coryphella lineata (Lovén, 1846) (additional source)
Galvinella Eliot, 1907 (status source)
Galvinella adarensis (Odhner, 1934) (basis of record)
Galvinella antarctica Eliot, 1907 (status source)
Galvinella glacialis Thiele, 1912 (status source)
Goniaeolididae Odhner, 1907 (status source)
Gulenia borealis (Odhner, 1922) accepted as Coryphella borealis Odhner, 1922 (status source)
Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Hantazuia Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuidae Korshunova, Fletcher & Martynov, 2025 (original description)
Himatina Thiele, 1931 accepted as Coryphella J. E. Gray, 1850 (status source)
Itaxia falklandica (Eliot, 1907) accepted as Coryphella falklandica Eliot, 1907 (additional source)
Janolina (status source)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) accepted as Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Microchlamylla gracilis (Alder & Hancock, 1844) accepted as Coryphella gracilis (Alder & Hancock, 1844) (status source)
Murmaniidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Notaeolidiidae Eliot, 1910 (additional source)
Notaeolidioidea Eliot, 1910 accepted as Fionoidea J. E. Gray, 1857 (basis of record)
Nudibranchus Martynov, 1998 (status source)
Aeolidina (status source)
Amphorina Quatrefages, 1844 (status source)
Amphorina amazighi (Tamsouri, Carmona, Moukrim & Cervera, 2015) (new combination reference)
Amphorina andra Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Amphorina farrani (Alder & Hancock, 1844) (additional source)
Amphorina linensis (García-Gómez, Cervera & F. J. Garcia, 1990) (additional source)
Amphorina pallida (Alder & Hancock, 1842) (additional source)
Amphorina viriola Korshunova, Malmberg, Prkić, Petani, Fletcher, Lundin & Martynov, 2020 (status source)
Annulorhina K. P. Rao, 1968 (status source)
Annulorhina mandapamensis K. P. Rao, 1968 (status source)
Apataidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Apataoidea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Fionoidea J. E. Gray, 1857 (basis of record)
Arminina (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Borealea nobilis (A. E. Verrill, 1880) accepted as Coryphella nobilis A. E. Verrill, 1880 (status source)
Borealea sanamyanae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Calmidae Iredale & O'Donoghue, 1923 accepted as Flabellinidae Bergh, 1889 (additional source)
Capellinia Trinchese, 1873 (status source)
Capellinia vittata (Alder & Hancock, 1842) (new combination reference)
Catriona aurantia (Alder & Hancock, 1842) (status source)
Chudidae Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo Korshunova, Fletcher & Martynov, 2025 (original description)
Chudo humanistica Korshunova, Fletcher & Martynov, 2025 (original description)
Chudoidea Korshunova, Fletcher & Martynov, 2025 (original description)
Corrupta Korshunova, Fletcher & Martynov, 2025 accepted as Coryphella J. E. Gray, 1850 (original description)
Coryphella alexanderi Ekimova, 2022 (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 (status source)
Coryphella borealis Odhner, 1922 (status source)
Coryphella browni Picton, 1980 (status source)
Coryphella californica Bergh, 1904 (status source)
Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 (status source)
Coryphella falklandica Eliot, 1907 (status source)
Coryphella fisheri MacFarland, 1966 accepted as Orienthella piunca (Er. Marcus, 1961) accepted as Coryphella piunca Er. Marcus, 1961 (source of synonymy)
Coryphella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) (additional source)
Coryphella lineata (Lovén, 1846) (status source)
Coryphella longicaudata O'Donoghue, 1922 (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella nobilis A. E. Verrill, 1880 (additional source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Coryphella trilineata O'Donoghue, 1921 (additional source)
Coryphella trophina (Bergh, 1890) (additional source)
Coryphella verta Ev. Marcus, 1970 (additional source)
Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025 (original description)
Cuthona Alder & Hancock, 1855 (additional source)
Cuthonellidae M. C. Miller, 1977 (additional source)
Cuthonidae Odhner, 1934 (additional source)
Dendronotina (status source)
Diaphoreolis flavovulta (MacFarland, 1966) (status source)
Diaphoreolis lagunae (O'Donoghue, 1926) (status source)
Diaphoreolis stipata (Alder & Hancock, 1843) (status source)
Diaphoreolis zvezda Korshunova, Fletcher, Bakken & Martynov, 2023 (status source)
Doridida (status source)
Dunga Eliot, 1902 (status source)
Embletonioidea Pruvot-Fol, 1954 (basis of record)
Eubranchidae Odhner, 1934 (additional source)
Eubranchopsis virginalis Baba, 1949 (status source)
Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchulus olivaceus (O'Donoghue, 1922) (status source)
Eubranchulus rupium (Møller, 1842) (status source)
Eubranchus Forbes, 1838 (additional source)
Eubranchus novik Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024 accepted as Eubranchulus novik (Grishina, Schepetov, Antokhina, Malaquias, Valdés & Ekimova, 2024) (status source)
Eubranchus rubropunctatus Edmunds, 1969 accepted as Produnga rubropunctata (Edmunds, 1969) (status source)
Favorinidae Bergh, 1889 accepted as Facelinidae Bergh, 1889 (additional source)
Fionoidea J. E. Gray, 1857 (basis of record)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Fjordia browni (Picton, 1980) accepted as Coryphella browni Picton, 1980 (status source)
Fjordia capensis (Thiele, 1925) accepted as Coryphella capensis Thiele, 1925 (status source)
Fjordia chriskaugei Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Fjordia insolita (García-Gómez & Cervera, 1989) accepted as Coryphella insolita (García-Gómez & Cervera, 1989) (status source)
Fjordia lineata (Lovén, 1846) accepted as Coryphella lineata (Lovén, 1846) (additional source)
Galvinella Eliot, 1907 (status source)
Galvinella adarensis (Odhner, 1934) (basis of record)
Galvinella antarctica Eliot, 1907 (status source)
Galvinella glacialis Thiele, 1912 (status source)
Goniaeolididae Odhner, 1907 (status source)
Gulenia borealis (Odhner, 1922) accepted as Coryphella borealis Odhner, 1922 (status source)
Gulenia monicae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Gulenia orjani Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (additional source)
Hantazuia Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025 (original description)
Hantazuidae Korshunova, Fletcher & Martynov, 2025 (original description)
Himatina Thiele, 1931 accepted as Coryphella J. E. Gray, 1850 (status source)
Itaxia falklandica (Eliot, 1907) accepted as Coryphella falklandica Eliot, 1907 (additional source)
Janolina (status source)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (status source)
Microchlamylla amabilis (Y. Hirano & Kuzirian, 1991) accepted as Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Microchlamylla gracilis (Alder & Hancock, 1844) accepted as Coryphella gracilis (Alder & Hancock, 1844) (status source)
Murmaniidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (additional source)
Notaeolidiidae Eliot, 1910 (additional source)
Notaeolidioidea Eliot, 1910 accepted as Fionoidea J. E. Gray, 1857 (basis of record)
Nudibranchus Martynov, 1998 (status source)
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Okinawa Island for Unidentia kiku Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Japanese part of the North Pacific Ocean for Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Okinawa Island for Unidentia kiku Korshunova, Fletcher & Martynov, 2025
Holotype KM1044, geounit Japan, identified as Hantazuia yugoikedai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1068, geounit Japan, identified as Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Holotype KM883, geounit Southern Ocean, identified as Chudo humanistica Korshunova, Fletcher & Martynov, 2025
Holotype KM954, verbatimGeounit USA, Washington Stat..., identified as Portorchardia candela Korshunova, Fletcher & Martynov, 2025
Holotype ZMMU Op-132.1, geounit Vietnam, identified as Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1059, geounit Okinawa Island, identified as Hantazuia imagawai Korshunova, Fletcher & Martynov, 2025
Holotype KM1068, geounit Japan, identified as Hantazuia kimotoi Korshunova, Fletcher & Martynov, 2025
Holotype KM883, geounit Southern Ocean, identified as Chudo humanistica Korshunova, Fletcher & Martynov, 2025
Holotype KM954, verbatimGeounit USA, Washington Stat..., identified as Portorchardia candela Korshunova, Fletcher & Martynov, 2025
Holotype ZMMU Op-132.1, geounit Vietnam, identified as Coryphellina iurmanovi Korshunova, Fletcher & Martynov, 2025
