WoRMS taxon details
Caryophylliidae Dana, 1846
135073 (urn:lsid:marinespecies.org:taxname:135073)
accepted
Family
Caryophylliadae Gray, 1847 · unaccepted > junior subjective synonym (misspelling)
Caryophylliinae Dana, 1846 · unaccepted > junior objective synonym
Parasmiliidae Alloiteau, 1952 † · unaccepted > junior homonym
Parasmiliidae Vaughan & Wells, 1943 † · unaccepted > junior subjective synonym
Parasmiliinae Vaughan & Wells, 1943 · unaccepted > junior subjective synonym
- Genus Angelismilia Reig Oriol, 1988 †
- Genus Anomocora Studer, 1878
- Genus Antillocyathus Wells, 1937 †
- Genus Asterosmilia Duncan, 1867 †
- Genus Aulocyathus Marenzeller, 1904
- Genus Bathycyathus Milne Edwards & Haime, 1848
- Genus Blastocyathus Reuss, 1865 †
- Genus Bourneotrochus Wells, 1984
- Genus Brachiatusmilia Alloiteau & Tissier, 1958 †
- Genus Caryophyllia Lamarck, 1801
- Genus Caryosmilia Wanner, 1902 †
- Genus Ceratotrochus Milne Edwards & Haime, 1848
- Genus Cereiophyllia Barta-Calmus, 1973 †
- Genus Clonosmilia Chevalier, 1963 †
- Genus Coenocyathus Milne Edwards & Haime, 1848
- Genus Coenosmilia Pourtalès, 1874
- Genus Colangia Pourtalès, 1871
- Genus Concentrotheca Cairns, 1979
- Genus Confluphyllia Cairns & Zibrowius, 1997
- Genus Conicosmilotrochus Turnšek, 1978 †
- Genus Conotrochus Seguenza, 1864
- Genus Crispatotrochus Tenison-Woods, 1878
- Genus Dasmosmilia Pourtalès, 1880
- Genus Dendrosmilia Milne Edwards & Haime, 1848 †
- Genus Desmophyllum Ehrenberg, 1834
- Genus Discocyathus Milne Edwards & Haime, 1848 †
- Genus Distolotrochus Kitahara & Cairns, 2021
- Genus Euafricana Ceccolini & Cianferoni, 2021
- Genus Faksephyllia Floris, 1972 †
- Genus Faustinotrochus Kitahara & Cairns, 2021
- Genus Frescocyathus Barta-Calmas, 1969 †
- Genus Ghirobocyathus Barta-Calmas, 1969 †
- Genus Goniocorella Yabe & Eguchi, 1932
- Genus Heterocyathus Milne Edwards & Haime, 1848
- Genus Hoplangia Gosse, 1860
- Genus Labyrinthocyathus Cairns, 1979
- Genus Leptocyathus Milne Edwards & Haime, 1850 †
- Genus Lochmaeotrochus Alcock, 1902
- Genus Lophosmilia Milne Edwards & Haime, 1848 †
- Genus Monohedotrochus Kitahara & Cairns, 2005
- Genus Multiplexcyathus Kitahara & Cairns, 2021
- Genus Nomlandia Durham & Barnard, 1952
- Genus Oxysmilia Duchassaing, 1870
- Genus Paraconotrochus Cairns & Parker, 1992
- Genus Paracyathus Milne Edwards & Haime, 1848
- Genus Paramontlivaltia Alloiteau, 1952 †
- Genus Parasmilia Milne Edwards & Haime, 1848 †
- Genus Phacelocyathus Cairns, 1979
- Genus Phyllangia Milne Edwards & Haime, 1848
- Genus Placocyathus Milne Edwards & Haime, 1848 †
- Genus Platycyathus de Fromentel, 1862 †
- Genus Polycyathus Duncan, 1876
- Genus Pourtalosmilia Duncan, 1884
- Genus Premocyathus Yabe & Eguchi, 1942
- Genus Rhizosmilia Cairns, 1978
- Genus Sclerhelia Milne Edwards & Haime, 1850
- Genus Smilotrochus Milne Edwards & Haime, 1851 †
- Genus Solenosmilia Duncan, 1873
- Genus Sphenotrochopsis Alloiteau & Tissier, 1958 †
- Genus Stephanosmilia de Fromentel, 1862 †
- Genus Stylocyathus d'Orbigny, 1849 †
- Genus Stylotrochus de Fromentel, 1862 †
- Genus Sympodangia Cairns & Zibrowius, 1997
- Genus Tethocyathus Kühn, 1933
- Genus Thalamophyllia Duchassaing, 1870
- Genus Thecocyathus Milne Edwards & Haime, 1848 †
- Genus Trochocyathus Milne Edwards & Haime, 1848
- Genus Vielicyathus Morycowa & Roniewicz, 1987 †
- Genus Vivesastraea Alloiteau & Tissier, 1958 †
- Genus Acanthocyathus Milne Edwards & Haime, 1848 accepted as Caryophyllia (Acanthocyathus) Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Actinosmilia d'Orbigny, 1849 † accepted as Lophosmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Africana Ocaña & Brito, 2015 accepted as Euafricana Ceccolini & Cianferoni, 2021 (unaccepted > junior homonym)
- Genus Agelecyathus Duncan, 1876 accepted as Polycyathus Duncan, 1876 (unaccepted > junior subjective synonym)
- Genus Amblocyathus d'Orbigny, 1849 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Anthophyllea Link, 1807 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Anthophyllum Schweigger, 1819 accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Anthrophyllum Schweigger, 1819 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym, misspelling)
- Genus Aplocyathus d'Orbigny, 1850 accepted as Trochocyathus (Aplocyathus) d'Orbigny, 1850 (unaccepted > superseded combination)
- Genus Blanfordia Duncan, 1880 † accepted as Platycyathus de Fromentel, 1862 † (unaccepted > junior subjective synonym)
- Genus Blastosmilia Duncan, 1878 accepted as Pourtalosmilia Duncan, 1884 (unaccepted > junior subjective synonym)
- Genus Brachycyathus Milne Edwards & Haime, 1848 † accepted as Discocyathus Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Brachytrochus Reuss, 1864 † accepted as Heterocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Brachytrochus Duncan, 1876 accepted as Heterocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym, homonym)
- Genus Brachytrossatus Strand, 1926 † accepted as Heterocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Carophyllia Lamarck, 1801 accepted as Caryophyllia (Caryophyllia) Lamarck, 1801 (unaccepted > misspelling - incorrect subsequent spelling)
- Subfamily Caryophylliinae Dana, 1846 accepted as Caryophylliidae Dana, 1846 (unaccepted > junior objective synonym)
- Genus Cenomanosmilia Alloiteau, 1952 † accepted as Lophosmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Ceratocyathus Seguenza, 1861 accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Ceratosmilia Alloiteau, 1957 † accepted as Smilotrochus Milne Edwards & Haime, 1851 † (unaccepted > junior subjective synonym)
- Genus Coelosmilia Milne Edwards & Haime, 1850 accepted as Anomocora Studer, 1878 (unaccepted > junior subjective synonym)
- Genus Cyathina Ehrenberg, 1834 accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Cyathoceras Moseley, 1880 accepted as Crispatotrochus Tenison-Woods, 1878 (unaccepted > junior subjective synonym)
- Genus Cyclicosmilia d'Orbigny, 1849 † accepted as Parasmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Cycliosmilia Milne Edwards & Haime, 1848 † accepted as Parasmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Cyclocyathus Milne Edwards & Haime, 1850 † accepted as Discocyathus Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Cylicosmilia Milne Edwards & Haime, 1848 accepted as Parasmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Cylindrocyathus Alloiteau, 1952 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Cyrtocyathus Alloiteau, 1958 † accepted as Trochocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Dasosmilia accepted as Dasmosmilia Pourtalès, 1880 (unaccepted > misspelling - incorrect subsequent spelling)
- Genus Dendrocyathus Alloiteau, 1952 † accepted as Dendrosmilia Milne Edwards & Haime, 1848 † (unaccepted > junior objective synonym)
- Genus Dungulia Oppenheim, 1930 † accepted as Smilotrochus Milne Edwards & Haime, 1851 † (unaccepted > junior subjective synonym)
- Genus Edwardsotrochus Chevalier, 1961 † accepted as Ceratotrochus (Edwardsotrochus) Chevalier, 1961 † (unaccepted > superseded combination)
- Genus Elasmosmilia Beauvais, 1961 † accepted as Trochocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Feddenia Duncan, 1880 accepted as Trochosmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Felixopsammia Filliozat, 1910 † accepted as Circophyllia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Fragilocyathus Yabe & Eguchi, 1932 accepted as Aulocyathus Marenzeller, 1904 (unaccepted > junior subjective synonym)
- Genus Gemmulatotrochus Duncan, 1878 accepted as Hoplangia Gosse, 1860 (unaccepted > junior subjective synonym)
- Genus Gemmulatrochus Duncan, 1878 † accepted as Hoplangia Gosse, 1860 (unaccepted > junior subjective synonym)
- Genus Goniocyathus Yabe & Eguchi, 1932 accepted as Dasmosmilia Pourtalès, 1880 (unaccepted > junior subjective synonym)
- Genus Lophelia Milne Edwards & Haime, 1849 accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Genus Lophohelia Milne Edwards & Haime, 1857 accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Genus Microcyathus Döderlein, 1913 accepted as Hoplangia Gosse, 1860 (unaccepted > junior subjective synonym)
- Genus Monocarya Lonsdale, 1850 † accepted as Parasmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Monticyathus Alloiteau & Tissier, 1958 † accepted as Bathycyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Odontocyathus Moseley, 1880 accepted as Stephanocyathus (Odontocyathus) Moseley, 1880 (unaccepted > junior subjective synonym)
- Subfamily Parasmiliinae Vaughan & Wells, 1943 accepted as Caryophylliidae Dana, 1846 (unaccepted > junior subjective synonym)
- Genus Parasmiliopsis Alloiteau, 1957 † accepted as Smilotrochus Milne Edwards & Haime, 1851 † (unaccepted > junior subjective synonym)
- Genus Paratrochocyathus Alloiteau, 1958 † accepted as Trochocyathus (Paratrochocyathis) Alloiteau, 1958 † (unaccepted > superseded combination, basionym)
- Genus Paterocyathus Duchassaing & Michelotti, 1860 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Phloeocyathus Alcock, 1902 accepted as Conotrochus Seguenza, 1864 (unaccepted > junior subjective synonym)
- Genus Platytrochopsis Sikharulidze, 1975 † accepted as Trochocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Plesiocaryophyllia Alloiteau, 1958 † accepted as Caryophyllia Lamarck, 1801 (unaccepted > junior subjective synonym)
- Genus Pleurocyathus Keferstein, 1859 accepted as Conocyathus d'Orbigny, 1849 (unaccepted > junior subjective synonym)
- Genus Pleurocyathus Moseley, 1881 † accepted as Conotrochus Seguenza, 1864 (unaccepted > junior subjective synonym)
- Genus Prototrochocyathus Kolosvary, 1959 † accepted as Parasmilia Milne Edwards & Haime, 1848 † (unaccepted > junior subjective synonym)
- Genus Psammocyathus Chevalier, 1963 † accepted as Crispatotrochus (Psammocyathus) Chevalier, 1963 † (unaccepted > superseded combination)
- Genus Psammosmilia de Fromentel, 1866 † accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Genus Pseudodesmophyllum Reig Oriol, 1997 † accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Genus Sabinotrochus Duncan, 1873 accepted as Stephanocyathus Seguenza, 1864 (unaccepted > junior subjective synonym)
- Genus Sclerohelia Milne Edwards & Haime, 1857 accepted as Sclerhelia Milne Edwards & Haime, 1850 (unaccepted > junior subjective synonym)
- Genus Scolangia Tenison-Woods, 1880 † accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Genus Soljania Pax, 1955 accepted as Hoplangia Gosse, 1860 (unaccepted > junior subjective synonym)
- Genus Spongiocyathus Folkeson, 1919 accepted as Heterocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Stellangia Duchassaing & Michelotti, 1860 accepted as Phyllangia Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Stephanoseris Milne Edwards & Haime, 1851 accepted as Heterocyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Stephanotrochus Moseley, 1880 accepted as Stephanocyathus Seguenza, 1864 (unaccepted > junior subjective synonym)
- Genus Steriphonotrochus Vaughan, 1900 † accepted as Bathycyathus Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Syndapas Lyman, 1859 accepted as Phyllangia Milne Edwards & Haime, 1848 (unaccepted > junior subjective synonym)
- Genus Thalassiotrochus Milne Edwards, 1861 accepted as Desmophyllum Ehrenberg, 1834 (unaccepted > junior subjective synonym)
- Subfamily Caryophylliinae Milne Edwards & Haime, 1857 (uncertain > taxon inquirendum)
- Genus Ellipsosmilia d'Orbigny, 1849 † (uncertain > nomen dubium)
- Genus Epitrochus de Fromentel, 1862 † (uncertain > nomen dubium)
marine, fresh, terrestrial
recent + fossil
Dana, J.D. (1846-1849). Zoophytes. United States Exploring Expedition during the years 1838-1842. <em>Lea and Blanchard, Philadelphia.</em> 7: 1-740, 61 pls. (1846: 1-120, 709-720; 1848: 121-708, 721-740; 1849: atlas pls. 1-61)., available online at https://www.biodiversitylibrary.org/page/18989497, http://www.sil.si.edu/digitalcollections/usexex/navigation/ScientificText/USExEx19_08select.cfm [details] 
Description This large family is usually divided into six sub-families, only one of which is hermatypic. The latter have phaceloid,...
Description This large family is usually divided into six sub-families, only one of which is hermatypic. The latter have phaceloid, meandroid or flabello-meandroid colonies with large, unperforated and widely spaced septa with little or no ornamentation. Corallite walls are of similar structure. The ahermatypic subfamilies are solitary or form phaceloid or dendroid colonies, usually with large-lobed septa and paliform lobes. All Cryophylliidae have a membraneous epitheca.
Related family is Flabellidae. (Veron, 1986 <57>). [details]
Related family is Flabellidae. (Veron, 1986 <57>). [details]
Hoeksema, B. W.; Cairns, S. (2025). World List of Scleractinia. Caryophylliidae Dana, 1846. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=135073 on 2025-04-27
Date
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Nomenclature
original description
Dana, J.D. (1846-1849). Zoophytes. United States Exploring Expedition during the years 1838-1842. <em>Lea and Blanchard, Philadelphia.</em> 7: 1-740, 61 pls. (1846: 1-120, 709-720; 1848: 121-708, 721-740; 1849: atlas pls. 1-61)., available online at https://www.biodiversitylibrary.org/page/18989497, http://www.sil.si.edu/digitalcollections/usexex/navigation/ScientificText/USExEx19_08select.cfm [details] 
original description (of Caryophylliinae Dana, 1846) Dana, J.D. (1846-1849). Zoophytes. United States Exploring Expedition during the years 1838-1842. <em>Lea and Blanchard, Philadelphia.</em> 7: 1-740, 61 pls. (1846: 1-120, 709-720; 1848: 121-708, 721-740; 1849: atlas pls. 1-61)., available online at https://www.biodiversitylibrary.org/page/18989497, http://www.sil.si.edu/digitalcollections/usexex/navigation/ScientificText/USExEx19_08select.cfm [details]
original description (of Parasmiliidae Alloiteau, 1952 †) Alloiteau J. (1952). Embranchement des Coelentérés. Madreporaires Post-Paleozoiques. <em>In: Piveteau J, ed. Traité de Paléontologie, Paris: Masson.</em> 539–684, pls. 1-10. [details]
original description (of Parasmiliinae Vaughan & Wells, 1943) Vaughan TW, Wells JW. (1943). Revision of the suborders, families and genera of the Scleractinia. <em>Special Papers of the Geological Society of America.</em> 44: 1-363. [details]
basis of record Cairns, S.D., Hoeksema, B.W., and J. van der Land, 2001. Scleractinia, <B><I>in</I></B>: Costello, M.J. <i>et al.</i> (Ed.) (2001). <i>European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,</i> 50: pp. 109-110 (look up in IMIS) [details]
original description (of Caryophylliinae Dana, 1846) Dana, J.D. (1846-1849). Zoophytes. United States Exploring Expedition during the years 1838-1842. <em>Lea and Blanchard, Philadelphia.</em> 7: 1-740, 61 pls. (1846: 1-120, 709-720; 1848: 121-708, 721-740; 1849: atlas pls. 1-61)., available online at https://www.biodiversitylibrary.org/page/18989497, http://www.sil.si.edu/digitalcollections/usexex/navigation/ScientificText/USExEx19_08select.cfm [details]
original description (of Parasmiliidae Alloiteau, 1952 †) Alloiteau J. (1952). Embranchement des Coelentérés. Madreporaires Post-Paleozoiques. <em>In: Piveteau J, ed. Traité de Paléontologie, Paris: Masson.</em> 539–684, pls. 1-10. [details]
original description (of Parasmiliinae Vaughan & Wells, 1943) Vaughan TW, Wells JW. (1943). Revision of the suborders, families and genera of the Scleractinia. <em>Special Papers of the Geological Society of America.</em> 44: 1-363. [details]
basis of record Cairns, S.D., Hoeksema, B.W., and J. van der Land, 2001. Scleractinia, <B><I>in</I></B>: Costello, M.J. <i>et al.</i> (Ed.) (2001). <i>European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,</i> 50: pp. 109-110 (look up in IMIS) [details]
Identification resource
identification resource
Cairns, S.D. & M. Kitahara. (2012). An illustrated key to the genera and subgenera of the Recent azooxanthellate Scleractinia (Cnidaria, Anthozoa), with an attached glossary. <em>ZooKeys.</em> 227: 1-47., available online at https://doi.org/10.3897/zookeys.227.3612 [details] Available for editors
[request]

Other
additional source
Hayward, P.J. & J.S. Ryland (Eds.). (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. <em>Clarendon Press: Oxford, UK.</em> 627 pp. (look up in IMIS) [details] Available for editors
[request]
additional source Seiblitz, I.G.L., C.F. Vaga, K.C.C. Capel, S.D. Cairns, J. Stolarski, A.M. Quattrini & M.V. Kitahara. (2022). Caryophylliids (Anthozoa, Scleractinia) and mitochondrial gene order: Insights from mitochondrial and nuclear phylogenomics. <em>Molecular Phylogenetics and Evolution.</em> 175: 107565., available online at https://doi.org/10.1016/j.ympev.2022.107565 [details] Available for editors
[request]

additional source Seiblitz, I.G.L., C.F. Vaga, K.C.C. Capel, S.D. Cairns, J. Stolarski, A.M. Quattrini & M.V. Kitahara. (2022). Caryophylliids (Anthozoa, Scleractinia) and mitochondrial gene order: Insights from mitochondrial and nuclear phylogenomics. <em>Molecular Phylogenetics and Evolution.</em> 175: 107565., available online at https://doi.org/10.1016/j.ympev.2022.107565 [details] Available for editors





From editor or global species database
Diagnosis Corallum solitary or colonial, free (polyp not investing entire corallum) or attached, most azooxanthellate, sometimes reproducing by transverse division; septa laminar, composed of one fan system of numerous closely-spaced, simple trabecular resulting in a smooth axial margin; wall septothecate or parathecate, sometimes covered with textura; endotheca and dissepiments sometimes present; synapticulae absent; septa arranged in various symmetries but usually hexameral; pali and columella present or absent; [details]Unreviewed
Description This large family is usually divided into six sub-families, only one of which is hermatypic. The latter have phaceloid, meandroid or flabello-meandroid colonies with large, unperforated and widely spaced septa with little or no ornamentation. Corallite walls are of similar structure. The ahermatypic subfamilies are solitary or form phaceloid or dendroid colonies, usually with large-lobed septa and paliform lobes. All Cryophylliidae have a membraneous epitheca.Related family is Flabellidae. (Veron, 1986 <57>). [details]