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Rushton-Mellor, S.K. (1994). The genus Argulus (Crustacea: Branchiura) in Africa: two new species, A. fryeri and A. gracilis, the previously undescribed male of A. brachypeltis Fryer and the identity of the male described as A. ambloplites Wilson. Systematic Parasitology. 28(1):23-31.
Rushton-Mellor, S.K. (1994). The genus Argulus (Crustacea: Branchiura) in Africa: two new species, A. fryeri and A. gracilis, the previously undescribed male of A. brachypeltis Fryer and the identity of the male described as A. ambloplites Wilson. Systematic Parasitology. 28(1):23-31.