Copepoda taxon details
Attheyella (Attheyella) crassa (Sars G.O., 1863)
716316 (urn:lsid:marinespecies.org:taxname:716316)
accepted
Species
Attheyella crassa (Sars G.O., 1863) · alternative representation
Attheyella crassa schmeili Gagern, 1938 · unaccepted
Attheyella spinosa Brady, 1880 · unaccepted (synonym)
Canthocamptus crassus Sars G.O., 1863 · unaccepted (synonym)
Canthocamptus horridus Fischer, 1860 · unaccepted
Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details] 
(of Canthocamptus crassus Sars G.O., 1863) Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details]
(of Canthocamptus crassus Sars G.O., 1863) Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details]
Type locality contained in Norway
type locality contained in Norway [details]
Taxonomic remark North-Eastern records in the Palearctic region possible belong to a different subspecies.
Taxonomic remark North-Eastern records in the Palearctic region possible belong to a different subspecies. [details]
Walter, T.C.; Boxshall, G. (2025). World of Copepods Database. Attheyella (Attheyella) crassa (Sars G.O., 1863). Accessed at: https://www.marinespecies.org/copepoda/aphia.php?p=taxdetails&id=716316 on 2026-08-06
Date
action
by
original description
Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details]
original description (of Attheyella crassa schmeili Gagern, 1938) Gagern, E. (1938). Harpacticoida des Harzes. [Harpacticoida of the Harz.]. <em>Zoologischer Anzeiger.</em> 124(7):175-182, figs. 1-12. (15-xi-1938). [details] Available for editors
[request]
original description (of Attheyella spinosa Brady, 1880) Brady, G.S. (1880). A monograph of the free and semi-parasitic Copepoda of the British Islands. Volume II. <em>Ray Society, London.</em> 2:1-182, pls. 34-82. (look up in IMIS) [details]
original description (of Canthocamptus crassus Sars G.O., 1863) Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details]
original description (of Canthocamptus horridus Fischer, 1860) Fischer, S. (1860). Beiträge zur Kenntniss der Entomostraceen. [Contributions to the knowledge of Entomostraca.]. <em>Abhandlungen der Bayerischen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Abteilung.</em> 8, Abt. 3:645-682, pls. 20-22., available online at https://www.biodiversitylibrary.org/page/16025063 [details]
additional source Apostolov, A. (1971). Copépodes Harpacticoïdes des sources de la montagne Strandja (Bulgarie). [Harpacticoid copepods from the springs of Strandja Mountain (Bulgaria).]. <em>Zoologischer Anzeiger.</em> 187(5-6):357-366, figs. 1-16. (xi/xii-1971). [details] Available for editors [request]
additional source Apostolov, A. & G.L. Pesce. (1989). Copepodes harpacticoides stygobies de Bulgarie. [Copepodes harpacticoides stygobies from Bulgaria.]. <em>Rivista di Idrobiologia.</em> 28(1-2):113-149. (vi-viii.1989). [details] Available for editors [request]
additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors [request]
additional source Chang, C.Y. & J.M. Lee. (2003). Taxonomy on freshwater canthocamptid harpacticoids from Korea II. Genus Attheyella. Korean Journal of Systematic Zoology 19(2): 189-201. [details] Available for editors [request]
additional source Chappuis, P.A. (1929). Révision du genre Canthocamptus Westwood (Note préliminaire). [Revision of the genus Canthocamptus Westwood (Preliminary note).]. <em>Buletinul Societatii de Stiinte din Cluj.</em> 4(2):41-50. (27-i-1929) [received Dec 1928]. [details] Available for editors [request]
additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors [request]
additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors [request]
additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors [request]
additional source Pesta, O. (1932). Krebstiere oder Crustacea. I: Ruderfüsser oder Copepoda. 3. Unterordnung: Harpacticoida (1. und 2. Hälfte). [Crustaceans or Crustacea. I: Copepods. 3. Suborder: Harpacticoida (1st and 2nd half)]. <em>In: Dahl, F. (ed.). Die Tierwelt Deutschlands und der Angrenzenden Meeresteile nach Ihren Merkmalen und nach Ihrer Lebensweise. Jena.</em> 24:1-164, figs. 1-72, 1-92. [details] Available for editors [request]
additional source Sarvala, J. (1977). The naupliar development of six species of freshwater harpacticoid Copepoda. Annales Zoologici Fennici 14(3):135-161, figs. 1-10. (15-ix-1977) [details] Available for editors [request]
additional source Shen, C.J. & T.H. Sung. (1965). On three new species and three new records of freshwater copepods from Sinkiang and Heilungkiang, China. <em>Acta Zootaxonomica Sinica.</em> 2(1):15-26, figs. 1-39. (i-1965, Chinese with English summary.). [details] Available for editors [request]
additional source Willems, K.A. (1989). Verspreiding, ecologie en gemeenschapsstructuur van benthische copepoden in het Delta-gebied en de Eems-Dollard (Nederland). [Distribution, ecology and community structure of benthic copepods in the Delta area and the Ems-Dollard (Netherlands)]. <em>Ph.D. Thesis, Universiteit Gent, Ghent, Belgium.</em> 2 vols:198 pp., available online at http://www.vliz.be/en/imis?module=dataset&dasid=5080) [details] Available for editors [request]
additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors [request]
additional source Sönmez, S. & S. Karaytug. (2019). Morphological and molecular contribution to the taxonomy of Attheyella (Attheyella) crassa (G. O. Sars, 1863) species complex (Harpacticoida: Canthocamptidae). Acta Zoologica Bulgarica, 71(1):9-16. [details] Available for editors [request]
additional source Caramujo, M.J. & M.J. Boavida. (2009). The practical identification of harpacticoids (Copepoda, Harpacticoida) in inland waters of central portugal for applied studies. <em>Crustaceana (Leiden).</em> 82(4):385-409., available online at https://doi.org/10.1163/156854009x404761 [details] Available for editors [request]
additional source Novikov, A.A., E.N. Abramova & R.M. Sabirov. (2020). Filogeniya semeystva Canthocamptidae (Copepoda, Harpacticoida) na osnove stroyeniya antenn i rotovykh konechnostey. [Phylogeny of Canthocamptidae (Copepoda, Harpacticoida) Based on the Morphology of Antennae and Mouthparts. ]. <em>Uchenye Zapiski Kazanskogo Universiteta. Seriya Estestvennye Nauki (Proceedings of Kazan University. Natural Sciences Series).</em> 162(4):629–647. [In Russian; English abstract]., available online at https://doi.org/10.26907/2542-064X.2020.4.629-647 [details] Available for editors [request]
additional source Kochanova, E., A. Nair, N. Sukhikh, R. Vainola & A. Husby. (2021). Patterns of Cryptic Diversity and Phylogeography in Four Freshwater Copepod Crustaceans in European Lakes. <em>Diversity-Basel.</em> 13(9):1-23. Sep 2021., available online at https://doi.org/10.3390/d13090448 [details] Available for editors [request]
additional source Dragos-Julian, C., C.A. Badaluta, R. Dumbrava, K.P. Battes & S. Iepure. (2025). Groundwater harpacticoids (Crustacea, Copepoda: Harpacticoida) in the karst areas of Apuseni mountains (northwestern Romania). <em>Travaux de l'Institut de Spéologie 'Émile Racovitza', Bucharest.</em> 64:113-143., available online at https://doi.org/10.59277/TISER.2025.04 [details] Available for editors [request]
additional source Kozel, P. (2018). Ecological evaluation of aquatic and terrestrial subterranean fauna in a karst cave. <em>Doctoral Dissertation, University of Nova Gorica, Nova Gorica, Slovenia.</em> 130 pp. [details] Available for editors [request]
original description (of Attheyella crassa schmeili Gagern, 1938) Gagern, E. (1938). Harpacticoida des Harzes. [Harpacticoida of the Harz.]. <em>Zoologischer Anzeiger.</em> 124(7):175-182, figs. 1-12. (15-xi-1938). [details] Available for editors
[request]original description (of Attheyella spinosa Brady, 1880) Brady, G.S. (1880). A monograph of the free and semi-parasitic Copepoda of the British Islands. Volume II. <em>Ray Society, London.</em> 2:1-182, pls. 34-82. (look up in IMIS) [details]
original description (of Canthocamptus crassus Sars G.O., 1863) Sars, G.O. (1863). Oversigt af de indenlandske Ferskvandcopepoder. [Overview of the domestic Freshwater Copepods.]. <em>Forhandlinger i Videnskabs-Selskabet i Christiana.</em> 1862:212-262., available online at https://www.biodiversitylibrary.org/page/44339019 [details]
original description (of Canthocamptus horridus Fischer, 1860) Fischer, S. (1860). Beiträge zur Kenntniss der Entomostraceen. [Contributions to the knowledge of Entomostraca.]. <em>Abhandlungen der Bayerischen Akademie der Wissenschaften. Mathematisch - Naturwissenschaftliche Abteilung.</em> 8, Abt. 3:645-682, pls. 20-22., available online at https://www.biodiversitylibrary.org/page/16025063 [details]
additional source Apostolov, A. (1971). Copépodes Harpacticoïdes des sources de la montagne Strandja (Bulgarie). [Harpacticoid copepods from the springs of Strandja Mountain (Bulgaria).]. <em>Zoologischer Anzeiger.</em> 187(5-6):357-366, figs. 1-16. (xi/xii-1971). [details] Available for editors
[request]additional source Apostolov, A. & G.L. Pesce. (1989). Copepodes harpacticoides stygobies de Bulgarie. [Copepodes harpacticoides stygobies from Bulgaria.]. <em>Rivista di Idrobiologia.</em> 28(1-2):113-149. (vi-viii.1989). [details] Available for editors
[request]additional source Bassamakov, I. (1971). Matériaux sur Harpacticoida (Copepoda) en Bulgarie. [Collections of Harpacticoida (Copepoda) in Bulgaria.]. <em>Godisnjak Muzeum v Plovdiv, Bulgaria.</em> 1965:9-19, figs. 1-8. (Bulgarian with French summary). [details] Available for editors
[request]additional source Chang, C.Y. & J.M. Lee. (2003). Taxonomy on freshwater canthocamptid harpacticoids from Korea II. Genus Attheyella. Korean Journal of Systematic Zoology 19(2): 189-201. [details] Available for editors
[request]additional source Chappuis, P.A. (1929). Révision du genre Canthocamptus Westwood (Note préliminaire). [Revision of the genus Canthocamptus Westwood (Preliminary note).]. <em>Buletinul Societatii de Stiinte din Cluj.</em> 4(2):41-50. (27-i-1929) [received Dec 1928]. [details] Available for editors
[request]additional source Damian-Georgescu, A. (1970). Copepoda Harpacticoida (forme de apa dulce). Fauna Republicii Socialiste Romania, Crustacea 4(11): 1-252, figs. 1-114. (14-vii-1970). [details] Available for editors
[request]additional source Ishida, T. (1994). Branchiopod and copepod crustaceans of Akan, northern Japan. Maeda Ippoen Foundation. Nature of Akan National Park 1993: 1241-1282. [In Japanese.] [details] Available for editors
[request]additional source Lang, K. (1931). Schwedische Süsswasser- und Moosharpacticiden. [Swedish freshwater and moss harpacticoids.]. <em>Arkiv för Zoologi.</em> (A)22(17):1-84, figs. 1-105. (19-iii-1931). [details] Available for editors
[request]additional source Pesta, O. (1932). Krebstiere oder Crustacea. I: Ruderfüsser oder Copepoda. 3. Unterordnung: Harpacticoida (1. und 2. Hälfte). [Crustaceans or Crustacea. I: Copepods. 3. Suborder: Harpacticoida (1st and 2nd half)]. <em>In: Dahl, F. (ed.). Die Tierwelt Deutschlands und der Angrenzenden Meeresteile nach Ihren Merkmalen und nach Ihrer Lebensweise. Jena.</em> 24:1-164, figs. 1-72, 1-92. [details] Available for editors
[request]additional source Sarvala, J. (1977). The naupliar development of six species of freshwater harpacticoid Copepoda. Annales Zoologici Fennici 14(3):135-161, figs. 1-10. (15-ix-1977) [details] Available for editors
[request]additional source Shen, C.J. & T.H. Sung. (1965). On three new species and three new records of freshwater copepods from Sinkiang and Heilungkiang, China. <em>Acta Zootaxonomica Sinica.</em> 2(1):15-26, figs. 1-39. (i-1965, Chinese with English summary.). [details] Available for editors
[request]additional source Willems, K.A. (1989). Verspreiding, ecologie en gemeenschapsstructuur van benthische copepoden in het Delta-gebied en de Eems-Dollard (Nederland). [Distribution, ecology and community structure of benthic copepods in the Delta area and the Ems-Dollard (Netherlands)]. <em>Ph.D. Thesis, Universiteit Gent, Ghent, Belgium.</em> 2 vols:198 pp., available online at http://www.vliz.be/en/imis?module=dataset&dasid=5080) [details] Available for editors
[request]additional source Chang, C.Y. (2009). Illustrated encyclopedia of fauna & flora of Korea. Vol. 42. Inland-water Copepoda. <em>Jeonghaeng-sa, Ministry of Education, Seoul, South Korea.</em> 42:1-687. [details] Available for editors
[request]additional source Sönmez, S. & S. Karaytug. (2019). Morphological and molecular contribution to the taxonomy of Attheyella (Attheyella) crassa (G. O. Sars, 1863) species complex (Harpacticoida: Canthocamptidae). Acta Zoologica Bulgarica, 71(1):9-16. [details] Available for editors
[request]additional source Caramujo, M.J. & M.J. Boavida. (2009). The practical identification of harpacticoids (Copepoda, Harpacticoida) in inland waters of central portugal for applied studies. <em>Crustaceana (Leiden).</em> 82(4):385-409., available online at https://doi.org/10.1163/156854009x404761 [details] Available for editors
[request]additional source Novikov, A.A., E.N. Abramova & R.M. Sabirov. (2020). Filogeniya semeystva Canthocamptidae (Copepoda, Harpacticoida) na osnove stroyeniya antenn i rotovykh konechnostey. [Phylogeny of Canthocamptidae (Copepoda, Harpacticoida) Based on the Morphology of Antennae and Mouthparts. ]. <em>Uchenye Zapiski Kazanskogo Universiteta. Seriya Estestvennye Nauki (Proceedings of Kazan University. Natural Sciences Series).</em> 162(4):629–647. [In Russian; English abstract]., available online at https://doi.org/10.26907/2542-064X.2020.4.629-647 [details] Available for editors
[request]additional source Kochanova, E., A. Nair, N. Sukhikh, R. Vainola & A. Husby. (2021). Patterns of Cryptic Diversity and Phylogeography in Four Freshwater Copepod Crustaceans in European Lakes. <em>Diversity-Basel.</em> 13(9):1-23. Sep 2021., available online at https://doi.org/10.3390/d13090448 [details] Available for editors
[request]additional source Dragos-Julian, C., C.A. Badaluta, R. Dumbrava, K.P. Battes & S. Iepure. (2025). Groundwater harpacticoids (Crustacea, Copepoda: Harpacticoida) in the karst areas of Apuseni mountains (northwestern Romania). <em>Travaux de l'Institut de Spéologie 'Émile Racovitza', Bucharest.</em> 64:113-143., available online at https://doi.org/10.59277/TISER.2025.04 [details] Available for editors
[request]additional source Kozel, P. (2018). Ecological evaluation of aquatic and terrestrial subterranean fauna in a karst cave. <em>Doctoral Dissertation, University of Nova Gorica, Nova Gorica, Slovenia.</em> 130 pp. [details] Available for editors
[request]
Present Inaccurate Introduced: alien Containing type locality
From editor or global species database
Additional information Phylogeography of the species based on populations from Russia, Europe and Fennoscandia [details]Habitat wells [details]
Habitat caves [details]
Habitat reed belt of a shallow lake [details]
Taxonomic remark North-Eastern records in the Palearctic region possible belong to a different subspecies. [details]
To Barcode of Life (35 barcodes) (from synonym Attheyella crassa (Sars G.O., 1863))
To Biodiversity Heritage Library (13 publications) (from synonym Attheyella crassa (Sars G.O., 1863))
To Biodiversity Heritage Library (26 publications) (from synonym Canthocamptus horridus Fischer, 1860)
To Biodiversity Heritage Library (43 publications) (from synonym Canthocamptus crassus Sars G.O., 1863)
To European Nucleotide Archive, ENA (Attheyella crassa) (from synonym Attheyella crassa (Sars G.O., 1863))
To GenBank (54 nucleotides; 34 proteins) (from synonym Canthocamptus crassus Sars G.O., 1863)
To GenBank (54 nucleotides; 34 proteins) (from synonym Attheyella crassa (Sars G.O., 1863))
To USNM Invertebrate Zoology Arthropoda Collection (1 record) (from synonym Attheyella crassa (Sars G.O., 1863))
To Biodiversity Heritage Library (13 publications) (from synonym Attheyella crassa (Sars G.O., 1863))
To Biodiversity Heritage Library (26 publications) (from synonym Canthocamptus horridus Fischer, 1860)
To Biodiversity Heritage Library (43 publications) (from synonym Canthocamptus crassus Sars G.O., 1863)
To European Nucleotide Archive, ENA (Attheyella crassa) (from synonym Attheyella crassa (Sars G.O., 1863))
To GenBank (54 nucleotides; 34 proteins) (from synonym Canthocamptus crassus Sars G.O., 1863)
To GenBank (54 nucleotides; 34 proteins) (from synonym Attheyella crassa (Sars G.O., 1863))
To USNM Invertebrate Zoology Arthropoda Collection (1 record) (from synonym Attheyella crassa (Sars G.O., 1863))
From editor or global species database
