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Blake, James A. (2025). New species and records of Scalibregmatidae (Annelida) from the Atlantic Ocean, Indian Ocean, Pacific Ocean, Southern Ocean, and adjacent seas. Megataxa. 16(1): 1–232.
507751
10.11646/megataxa.16.1.1 [view]
urn:lsid:zoobank.org:pub:0AC91FF7-0A79-47EC-95C6-0FC661AE68FF [view]
Blake, James A.
2025
New species and records of Scalibregmatidae (Annelida) from the Atlantic Ocean, Indian Ocean, Pacific Ocean, Southern Ocean, and adjacent seas
Megataxa
16(1): 1–232
Publication
Annelidabase, open access
A large collection of scalibregmatid polychaetes has been assembled from offshore monitoring and reconnaissance surveys conducted in the Western North Atlantic Ocean, Gulf of Mexico, Eastern North Pacific Ocean, Southern Ocean, and South China Sea. The materials from these projects and others from several museum collections have yielded specimens from ten genera and 76 species, of which 54 are new to science. The type species of the genera Asclerocheilus, Axiokebuita, Oligobregma, Scalibregmella, Sclerobregma, and Mucibregma are redescribed either from type material and/or specimens from near the type locality. The type-species of Axiokebuita (A. millsi) is revalidated and redescribed based on new material off eastern North America. Scalibregma brevicaudum Verrill, 1873 is revalidated based on the holotype from Long Island Sound and numerous new collections from northeastern North America. The ten genera treated are as follows: Asclerocheilus 16 species with 12 new to science; Axiokebuita six species with three new to science; Hyboscolex six species with five new to science; Oligobregma 14 species with 11 new to science; Parasclerocheilus one new species; Pseudoscalibregma eight species with five new to science; Scalibregma 19 species with 15 new to science; Scalibregmella two species with one new to science; Sclerobregma two species with one new to science, and Mucibregma, one species. Most new species are from offshore or deep-water habitats and, except in areas with a history of extensive monitoring, are represented by relatively few individuals. Known species of each genus with five or more species are tabulated, compared, and reviewed. The depth intervals of all scalibregmatids are tabulated in order of increasing depth. Lists of deep-water scalibregmatids are updated. Of 141 known species of Scalibregmatidae, 73 or 52 % occur in depths of 1000 m or greater. Of the 54 new species included in this report, nine occur from the intertidal to 100 m; four occur from 100 to 500 m; six occur from 500 to 1000 m; and 34 species or 63% occur in depths greater than 1000 m, with 12 of these ranging into abyssal depths of ~3000 m or deeper. The status of Mucibregma spinosa Fauchald & Hancock, 1981 is reviewed and, although poorly preserved and damaged, is confirmed as a species of Scalibregmatidae, albeit unique with acicular spines on all parapodia. The holotype of the enigmatic deep-water genus and species Neolipobranchius glabrus Hartman & Fauchald, 1971 is demonstrated to be an indeterminate juvenile of the genus Travisia and, with the genus becoming a junior synonym of Travisia, the species is declared a species inquirenda.
Systematics, Taxonomy
RIS (EndNote, Reference Manager, ProCite, RefWorks)
BibTex (BibDesk, LaTeX)
Date
action
by
2025-04-16 05:43:42Z
created

Asclerocheilus Ashworth, 1901 (taxonomy source)
Asclerocheilus alatus Blake, 2025 (original description)
Asclerocheilus aurospinosus Blake, 2025 (original description)
Asclerocheilus beringianus Uschakov, 1955 (taxonomy source)
Asclerocheilus caroliniensis Blake, 2025 (original description)
Asclerocheilus intermedius (Saint-Joseph, 1894) (redescription)
Asclerocheilus jonesi Blake, 2025 (original description)
Asclerocheilus nathanae Blake, 2025 (original description)
Asclerocheilus neubertae Blake, 2025 (original description)
Asclerocheilus pseudocollare (Schüller & Hilbig, 2007) accepted as Asclerocheilus pseudocollaris (Schüller & Hilbig, 2007) (new combination reference)
Asclerocheilus triangularis Blake, 2025 (original description)
Axiokebuita Pocklington & Fournier, 1987 (taxonomy source)
Axiokebuita australis Blake, 2023 (additional source)
Axiokebuita millsi Pocklington & Fournier, 1987 (status source)
Axiokebuita minuta (Hartman, 1967) (taxonomy source)
Axiokebuita shackletoni Blake, 2025 (original description)
Axiokebuita verenae Blake, 2025 (original description)
Hyboscolex Schmarda, 1861 (taxonomy source)
Hyboscolex antarcticus Blake, 2025 (original description)
Hyboscolex farallonensis Blake, 2025 (original description)
Hyboscolex reesi Blake, 2025 (original description)
Mucibregma Fauchald & Hancock, 1981 (taxonomy source)
Mucibregma spinosa Fauchald & Hancock, 1981 accepted as Mucibregma spinosum Fauchald & Hancock, 1981 (redescription)
Neolipobranchius Hartman & Fauchald, 1971 accepted as Travisia Johnston, 1840 (status source)
Oligobregma Kudenov & Blake, 1978 (taxonomy source)
Oligobregma aciculata (Hartman, 1965) accepted as Oligobregma aciculatum (Hartman, 1965) (taxonomy source)
Oligobregma acsarum Blake, 2025 (original description)
Oligobregma acutaspinosum Blake, 2025 (original description)
Oligobregma blakei Schüller & Hilbig, 2007 accepted as Asclerocheilus pseudocollare (Schüller & Hilbig, 2007) accepted as Asclerocheilus pseudocollaris (Schüller & Hilbig, 2007) (source of synonymy)
Oligobregma brychiatum Blake, 2025 (original description)
Oligobregma chromatum Blake, 2025 (original description)
Oligobregma emmae Blake, 2025 (original description)
Oligobregma escanabis Blake, 2025 (original description)
Oligobregma hampsoni Blake, 2025 (original description)
Oligobregma menziesi Blake, 2025 (original description)
Oligobregma mucronata Blake, 2015 accepted as Oligobregma mucronatum Blake, 2015 (taxonomy source)
Oligobregma sphaeracirratum Blake, 2025 (original description)
Oligobregma weddelliensis Blake, 2025 accepted as Oligobregma weddelliense Blake, 2025 (original description)
Oligobregma whaleyi Wiklund, Neal, Glover, Drennan, Rabone & Dahlgren, 2019 (taxonomy source)
Oligobregma williamsae Blake, 2025 (original description)
Pseudoscalibregma Ashworth, 1901 (taxonomy source)
Pseudoscalibregma biannulatum Blake, 2025 (original description)
Pseudoscalibregma bransfieldia (Hartman, 1967) (taxonomy source)
Pseudoscalibregma gracilis Blake, 2025 accepted as Pseudoscalibregma gracile Blake, 2025 (original description)
Pseudoscalibregma larae Blake, 2025 (original description)
Pseudoscalibregma nybakkeni Blake, 2025 (original description)
Pseudoscalibregma palmeri Blake, 2015 (taxonomy source)
Pseudoscalibregma parvum (Hansen, 1879) (taxonomy source)
Pseudoscalibregma pigmentum Blake, 2025 (original description)
Scalibregma antennatum Blake, 2025 (original description)
Scalibregma aubreyi Blake, 2025 (original description)
Scalibregma australis Blake, 2015 accepted as Scalibregma australe Blake, 2015 (taxonomy source)
Scalibregma brevicaudum [auct. misspelling] accepted as Scalibregma brevicauda Verrill, 1873 (taxonomy source)
Scalibregma californicum Blake, 2000 (taxonomy source)
Scalibregma cornutum Blake, 2025 (original description)
Scalibregma fimbriatum Blake, 2025 (original description)
Scalibregma hadrostomium Blake, 2025 (original description)
Scalibregma inflatum Rathke, 1843 (taxonomy source)
Scalibregma lydoniensis Blake, 2025 accepted as Scalibregma lydoniense Blake, 2025 (original description)
Scalibregma profundum Blake, 2025 (original description)
Scalibregma rhoadsi Blake, 2025 (original description)
Scalibregmatidae Malmgren, 1867 (taxonomy source)
Scalibregmella Hartman & Fauchald, 1971 (taxonomy source)
Scalibregmella antennata Hartman & Fauchald, 1971 (redescription)
Scalibregmella pacifica Blake, 2025 (original description)
Sclerobregma branchiatum Hartman, 1965 (taxonomy source)
Sclerobregma multiaciculatum Blake, 2025 (original description)
Holotype LACM LACM-AHF Poly 13574., geounit Peru-Chile Trench, identified as Pseudoscalibregma biannulatum Blake, 2025
Holotype MCZ IZ 171910, geounit Gulf of Mexico, identified as Hyboscolex reesi Blake, 2025
Holotype USNM 1741704, geounit North West Atlantic, identified as Oligobregma sphaeracirratum Blake, 2025
Holotype USNM 1741779., geounit West North Atlantic, identified as Scalibregma fimbriatum Blake, 2025
Holotype LACM AHF-Poly 13308, geounit Farallon Islands, identified as Pseudoscalibregma nybakkeni Blake, 2025
Holotype LACM LACM-AHF Poly 13310, geounit West North Atlantic, identified as Oligobregma williamsae Blake, 2025
Holotype LACM LACM-AHF Poly 13557, geounit Clarion Clipperton Zone, identified as Scalibregmella pacifica Blake, 2025
Holotype LACM LACM-AHF Poly 13570, geounit Mozambique Channel, identified as Scalibregma cornutum Blake, 2025
Holotype LACM LACM-AHF Poly 13573, geounit Peru-Chile Trench, identified as Scalibregma profundum Blake, 2025
Holotype LACM LACM-AHF Poly 13575, geounit Peru-Chile Trench, identified as Oligobregma menziesi Blake, 2025
Holotype LACM LACM-AHF Poly 14834., geounit Farallon Islands, identified as Hyboscolex farallonensis Blake, 2025
Holotype LACM LACM-AHF Poly 14838, geounit Farallon Islands, identified as Scalibregma antennatum Blake, 2025
Holotype LACM LACM-AHF Poly 14841, geounit Farallon Islands, identified as Scalibregma hadrostomium Blake, 2025
Holotype MCZ IZ 171914, geounit Borneo, identified as Oligobregma chromatum Blake, 2025
Holotype MCZ IZ 171915, geounit Borneo, identified as Oligobregma emmae Blake, 2025
Holotype MCZ IZ 171922, geounit Gorda Ridges, identified as Oligobregma escanabis Blake, 2025
Holotype MCZ IZ 171932, geounit Brunei, identified as Pseudoscalibregma pigmentum Blake, 2025
Holotype MCZ IZ 171936, geounit Brunei, identified as Scalibregma aubreyi Blake, 2025
Holotype MCZ IZ 172024, geounit Brunei, identified as Sclerobregma multiaciculatum Blake, 2025
Holotype USNM 1741695, geounit North Carolina, identified as Oligobregma acsarum Blake, 2025
Holotype USNM 1741700, geounit South Carolina, identified as Oligobregma acutaspinosum Blake, 2025
Holotype USNM 1741703., geounit North Atlantic, identified as Oligobregma hampsoni Blake, 2025
Holotype USNM 1741705., geounit North Carolina, identified as Pseudoscalibregma larae Blake, 2025
Holotype USNM 1741809, geounit Georges Bank, identified as Scalibregma lydoniensis Blake, 2025
Holotype USNM 741813, geounit Cape Hatteras, identified as Scalibregma rhoadsi Blake, 2025
Holotype ZMH ZMH-ANN-030785, geounit Scotia Sea, identified as Oligobregma brychiatum Blake, 2025
Holotype ZMH ZMH-ANN-030790., geounit Weddell Sea, identified as Oligobregma weddelliensis Blake, 2025
Holotype ZMH ZMH-ANN-030796, geounit Scotia Sea, identified as Pseudoscalibregma gracilis Blake, 2025
 Diagnosis

[Diagnosis of Blake, 2025] Body elongate, arenicoliform. Prostomium with a pair or lateral or frontal horns. ... [details]

 Diagnosis

[Original diagnosis of Pocklington & Fournier (1987)] Body fusiform, 20-30 segments; prostomium triangular with ... [details]

 Diagnosis

[Diagnosis of Blake (2025)] Body elongate, arenicoliform. Prostomium with frontal horns; eyespots present or ... [details]

 Diagnosis

(Emended Blake, 2025: 202). Body elongate, arenicoliform, expanded anteriorly. Prostomium T-shaped with lateral ... [details]

 Diagnosis

Body elongate, arenicoliform. Prostomium T-shaped with frontal horns. Peristomium a single ring, encompassing ... [details]

 Diagnosis

"Body elongate, arenicoliform. Prostomium T-shaped with lateral horns. Peristomium achaetous, surrounding ... [details]

 Distribution

According to Blake (2025) Axiokebuita minuta is widespread in the Southern Ocean in continental shelf and slope ... [details]

 Distribution

Blake (2025) lists numerous reports of Hyboscolex pacificus since the original description by Moore (1909) as ... [details]

 Editor's comment

Lipobranchius is currently monotypic, containing only Lipobranchius jeffreysii (McIntosh, 1869) . However, it also ... [details]

 Etymology

Asclerocheilus nathanae is named for is named for Ms. Desiree Nathan, Kuala Lumpur, Malaysia, who was overall ... [details]

 Etymology

Axiokebuita verenae is named after Dr. Verena Tunnicliffe, original collector of the species. [details]

 Etymology

Hyboscolex ehlersi is named for Polychaeta taxonomist Professor Ernst Heinrich Ehlers (1835–1925) [details]

 Etymology

Hyboscolex farallonensis is named after the proximity of the offshore type locality to the Farallon Islands, off ... [details]

 Etymology

Hyboscolex harrisae is named for Leslie Harris, joint collector of the specimens, and Collections Manager at LACM. [details]

 Etymology

Hyboscolex reesi is named for Dr. Bryant Eugene Rees (1912–1985), Professor of Biology at California State ... [details]

 Etymology

Oligobregma acsarum is named from "ACSAR, the acronym used by the project team for the Atlantic Continental Slope ... [details]

 Etymology

Oligobregma acutaspinosum is named from the Latin acutus, for pointed and the Latin spina, for thorn in reference ... [details]

 Etymology

Oligobregma brychiatum is named for "Greek, brychios meaning “from the deep” in reference to the great depths ... [details]

 Etymology

Oligobregma chromatum is named from the Greek ‘chroma’, for color of the skin or surface, in reference to the ... [details]

 Etymology

Oligobregma emmae is named after the R/V Emma, the research vessel used on the survey to collect samples from off ... [details]

 Etymology

Oligobregma escanabis is named for the Escanaba Trough, where collected. It is not stated how 'escanabis' is ... [details]

 Etymology

Oligobregma hampsoni is named after George W. Hampson, former Research Scientist at the Woods Hole Oceanographic ... [details]

 Etymology

Oligobregma menziesi is named after Dr. Robert J. Menzies, Chief Scientist on several of the SEPBOB voyages. [details]

 Etymology

Oligobregma sphaeracirratum is named for the large spherical dorsal cirri on the posterior parapodia, intended as a ... [details]

 Etymology

Oligobregma weddelliensis Blake, 2025 is named after the Weddell Sea, Antarctica, where collected. Note that the ... [details]

 Etymology

Oligobregma williamsae is named after Ms. Isabelle P. (Izzie) Williams who worked with the late Dr. Howard L. ... [details]

 Etymology

Parasclerocheilus fonsecae is named after the collector of the holotype, the late Ana C. Fonseca-Escalante, Costa ... [details]

 Etymology

Pseudoscalibregma biannulatum is named as a compound formed from bi, Latin for two and annulus, Latin for ring, in ... [details]

 Etymology

Pseudoscalibregma gracilis is named from the Latin gracilis, for slender or thin in reference to the unusually ... [details]

 Etymology

Pseudoscalibregma larae is named after Ms. Lara (Luzak) Parisi, graduate assistant in 2007 who investigated the ... [details]

 Etymology

Pseudoscalibregma nybakkeni is named after Dr. James W. Nybakken (1936–2009), former Professor of Marine Biology ... [details]

 Etymology

Pseudoscalibregma pigmentum is named for Latin 'pigmentum' color or paint, after the pigmented posterior glands. ... [details]

 Etymology

Scalibregma aggregatum is named for adjectival Latin aggregatus -a -um meaning "collected' or 'clustered' ... [details]

 Etymology

Scalibregma antennatum is named for Latin, antenna, a sensory appendage on the head in reference to the elongate, ... [details]

 Etymology

Scalibregma aubreyi is named after the late Mr. Stephen B. Aubrey, oceanographic field scientist on the sampling ... [details]

 Etymology

Scalibregma beaufortensis is named for the Beaufort Sea in the Arctic Ocean where it was collected. For neuter ... [details]

 Etymology

Not stated. The noun phrase 'brevicauda' is Latin for short tail. 'Cauda' (tail) is a noun and nouns or compound ... [details]

 Etymology

Scalibregma chilensis is named for its country of origin. For neuter genus gender agreement the correct adjectival ... [details]

 Etymology

Scalibregma cornutum is named for the distinctive laterally directed horns on the anterior margin of prostomium, a ... [details]

 Etymology

Scalibregma fimbriatum is named for the unusual double row of lobes on the anterior margin of the two peristomial ... [details]

 Etymology

Scalibregma hadrostomium is named for its enlarged and inflated peristomium, a neuter compound Latin adjective ... [details]

 Etymology

Scalibregma lydoniensis is named after Lydonia Canyon. For neuter genus gender agreement the correct adjectival ... [details]

 Etymology

Scalibregma profundum is named for the abyssal depths of the type locality from Latin neuter adjective 'profundus'. [details]

 Etymology

Scalibregma rhoadsi is named after named for Dr. Donald C. Rhoads, benthic ecologist and part of the Cape Hatteras ... [details]

 Etymology

Scalibregma ridleyensis is named for Ridley Island near where collected. For neuter genus gender agreement the ... [details]

 Etymology

Scalibregma sahlingi is named for the collector of the species, Dr. Heiko Sahling (1969–2018), marine ecologist ... [details]

 Etymology

Scalibregma therouxi is named after Dr. Roger B. Theroux (1935–2008), former biologist with the U.S. National ... [details]

 Etymology

Scalibregma velatum is named for the annular rings of the peristomium and chaetiger 1 that together form an ... [details]

 Etymology

Scalibregmella pacifica is named for the ocean in which it was found. [details]

 Etymology

Sclerobregma multiaciculatum is named for the multiple kinds of anterior acicular spines present, a compound ... [details]

 Grammatical gender

Cheilus' (cheilos) could be neuter, but Grube treated it as masculine, as did following authors and the authors of ... [details]

 Holotype

Blake (2025: 162) revalidated Scalibregma brevicauda (as 'brevicaudum') and reported the probable holotype (from ... [details]

 Identification

Blake (2025: 25) states that, alfthough reported widely "It is clear that most, if not all, of the non-Arctic ... [details]

 Spelling

Asclerocheilus has been treated as masculine, therefore as a masculine genus the name when recombined should be ... [details]

 Spelling

Oligobregma aciculata should be O. aciculatum (q.v) for grammatical agreement of specific epithet with a neuter ... [details]

 Spelling

Note that the author published the species-group name as 'wedelliensis' instead of the correct neuter 'wedelliense' ... [details]

 Spelling

The author's published spelling of 'gracilis' should be corrected to 'gracile'. For a neuter genus gender ... [details]

 Spelling

As Scalibregma is a neuter gender genus it is mandatory to correct the adjectival spelling to australe as ... [details]

 Spelling

Note that the author published the species-group name as 'beaufortensis' instead of the correct neuter ... [details]

 Spelling

Blake 2025:162) revalidated Scalibregma brevicauda from long-existing synonymy with S. inflatum and elected to ... [details]

 Spelling

Note that the author published the species-group name as 'chilensis' instead of the correct neuter 'chilense'. For ... [details]

 Spelling

Note that the author published the species-group name as 'lydoniensis' instead of the correct neuter 'lydoniense'. ... [details]

 Spelling

Note that the author published the species-group name as 'ridleyensis' instead of the correct neuter 'ridleyense'. ... [details]

 Spelling

Hartman (1965) correctly created a neuter adjectival species-group name of branchiatum, but subsequently (e.g. ... [details]

 Spelling

Hartman (1965) correctly creates an neuter adjectival species-group name of branchiatum, but subsequently (e.g. ... [details]

 Status

The holotype of N. glabrus is not identifiable to genus or species and is a juvenile (fide Blake 2015: [as spelling ... [details]

 Synonymy

Blake (2025: 43) as first revisor placed O. blakei as synonym, based on a juvenile described from same station as ... [details]

 Taxonomy

Kudenov (1985:339), examined the only specimen and suggested Mucibregma spinosa might be a member of ... [details]

 Type locality

Stephens Passage, near Juneau, Alaska, NE Pacific Ocean, 58.3040°N, 134.7095°W, ca. 50 m, holotype CASIZ 165339 [details]

 Type locality

west of Farallon Islands, Continental slope off northern California, 37.75°N, 123.583°W, 2850 m, holotype ... [details]

 Type locality

continental slope off Durban, Natal, South Africa, Indian Ocean, 30.200°S, 32.017°E, 1360 m, holotype LACM-AHF ... [details]

 Type locality

W of Nelson Island, Scotia Sea, off Antarctic Peninsula, 63.225°S, 64.613°W, 2079 m, holotype ZMHANN- 030779 [details]

 Type locality

off Charleston, South Carolina, North America, 32.2°N, 76.704°W, 1991 m, holotype USNM 1741686 [details]

 Type locality

rock scraping, Santa Maria Basin off Purisima Point, California, 34.759ºN, 129.837ºW to 34.765ºN, 129.829ºW, ... [details]

 Type locality

south of Pioneer Canyon, off San Francisco, California, North America, 37.225ºN, 123.125ºW, 1020 m, holotype ... [details]

 Type locality

Matterhorn Platform Survey, off Louisiana, Gulf of Mexico, 28.742°N, 88.805°W, 921 m, holotype MCZ IZ 171905. [details]

 Type locality

E of Fuerte Bulnes, Strait of Magellan, Tiera del Fuego, 53.633°S, 70.842°W, 249–392 m, holotype LACM-AHF Poly ... [details]

 Type locality

off Brunei, Island of Borneo, South China Sea, 5.523°N, 113.705°E, 1947 m, holotype MCZ IZ 171906 [details]

 Type locality

West of Farallon Islands Continental slope off northern California, 37.632°N, 123.448°W, 2670 m, holotype ... [details]

 Type locality

West of Farallon Islands, Continental slope off northern California, 37.717°N, 123.4670°W, 2250 m, ... [details]

 Type locality

Las Gemelas Seamount, near Isla del Coco, off Costa Rica, East Pacific Ocean, 4.9871°N, 87.4433°W, 243 m, ... [details]

 Type locality

Abyssal plain off Cape Norvegia, Weddell Sea, Antarctica, 70.640831°S, 14.714330°W, 4928 m, holotype ZMH-ANN-030783. [details]

 Type locality

Slurp sample on basalt, HOV Pisces IV, Hell vent, Southern Axial Seamount, Juan de Fuca Ridge, 45.932°N, ... [details]

 Type locality

Abyssal plain West of Antarctic Peninsula, Weddell Sea, 65.2700°S, 48.06350°W, 4069 m, holotype ZMH-ANN-030784. [details]

 Type locality

Rocks in sand, Western side Isla Quenu, Punta Pinto, Golfo de Ancud, Chile, 41.821°S, 73.171°W, intertidal, ... [details]

 Type locality

West of Farallon Islands, West of San Francisco, Continental slope off northern California, 37.7°N, 123.45°W, ... [details]

 Type locality

Off Tjarno Marine Station, Kosterfjord, Skagerrak, Sweden, 58.882°N, 11.095°E to 58.882°N, 11.098°E, 50–100 ... [details]

 Type locality

Matterhorn Platform Survey, off Louisiana, Northern Gulf of Mexico, 28.742°N, 88.831°W, 850 m, holotype MCZ IZ 171910. [details]

 Type locality

off Cape Fear, North Carolina, North America Atlantic coast, South ACSAR Program, 32.921°N, 75.833°W, 3006 m, ... [details]

 Type locality

Offshore of Charleston, South Carolina, 32.395°N, 77.018°W, 800 m, holotype USNM 1741700. [details]

 Type locality

Scotia Sea, Antarctica, 60.449167°S, 56.082667°W, 3957 m, holotype ZMH-ANN-030785. [details]

 Type locality

Offshore of Brunei, Island of Borneo, 5.517°N, 113.608°E, 1999 m, holotype MCZ IZ 171914. [details]

 Type locality

Offshore of Brunei, Island of Borneo, 5.36°N, 113.821°E, 1325 m, holotype MCZ IZ 171915 [details]

 Type locality

HOV Alvin Dive 2035, Gorda Ridge, Escanaba Trough, N Pacific, 41.007°N, 127.293°W, 3305 m, box core B, holotype ... [details]

 Type locality

Offshore of New England, North America Atlantic coast, 40.954°N, 66.230°W, 2103 m, holotype USNM 1741703. [details]

 Type locality

Milne-Edwards Deep, Peru-Chile Trench, offshore of Peru, 08.7°S, 80.667°W, 3167–3318 m, holotype LACM-AHF Poly ... [details]

 Type locality

Offshore New England, NW Atlantic, 40.085°N, 67.497°W, 2071 m, holotype USNM 1741704. [details]

 Type locality

Weddell Sea, E of Antarctic Peninsula, 65.3108 S, 51.5152 W, 3065 m, holotype ZMH-ANN-030790. [details]

 Type locality

Off New Jersey, Gay-Head Bermuda Transect, 38.267°N, 71.783°W, 2864 m, holotype LACM-AHF Poly 13310. [details]

 Type locality

Coast of Punta Islotes, Golfo Dulce, Costa Rica, Pacific coast, 8.7327 N, 83.3979 W, intertidal, holotype MCZ IZ 62651. [details]

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