1. Subfamily Pinnotherelinae Alcock, 1900
2. Subfamily Varuninae H. Milne Edwards, 1853
   1. Genus Brachynotus De Haan, 1833
   2. Genus Eriocheir De Haan, 1835
   3. Genus Hemigrapsus Dana, 1851

marine, brackish, fresh, terrestrial

Milne Edwards, H. (1853-1854). Mémoire sur la famille des ocypodiens. <em>Annales des Sciences Naturelles, 3rd series.</em> 20: 163-228, pls. 4-13 [163-192, pls 4-5 published Dec 1853; 193-228, pls 6-13 published Jan 1854; see Clark, 2025). [details] 

DecaNet eds. (2025). DecaNet. Varunidae H. Milne Edwards, 1853. Accessed through: Costello, M. J.; Ahyong, S.; Bieler, R.; Boudouresque, C.; Desiderato, A.; Downey, R.; Galil, B. S.; Gollasch, S.; Hutchings, P.; Kamburska, L.; Katsanevakis, S.; Kupriyanova, E.; Lejeusne, C.; Ma, K. C. K.; Marchini, A.; Occhipinti, A.; Pagad, S.; Pino, L.; Poore, G. C. B.; Rewicz, T.; Rius, M.; Robinson, T. B.; Sobczyk, R.; Stepien, A.; Turon, X.; Valls Domedel, G.; Verleye, T.; Vieira, L. M.; Willan, R. C.; Zhan, A. (2025) World Register of Introduced Marine Species (WRiMS) at: https://www.marinespecies.org/Introduced/aphia.php?p=taxdetails&id=439140 on 2026-04-17

Costello, M. J.; Ahyong, S.; Bieler, R.; Boudouresque, C.; Desiderato, A.; Downey, R.; Galil, B. S.; Gollasch, S.; Hutchings, P.; Kamburska, L.; Katsanevakis, S.; Kupriyanova, E.; Lejeusne, C.; Ma, K. C. K.; Marchini, A.; Occhipinti, A.; Pagad, S.; Pino, L.; Poore, G. C. B.; Rewicz, T.; Rius, M.; Robinson, T. B.; Sobczyk, R.; Stępień, A.; Turon, X.; Valls Domedel, G.; Verleye, T.; Vieira, L. M.; Willan, R. C.; Zhan, A. (2026). World Register of Introduced Marine Species (WRiMS). Varunidae H. Milne Edwards, 1853. Accessed at: https://www.marinespecies.org/introduced/aphia.php?p=taxdetails&id=439140 on 2026-04-17

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