Hookabe, N.; Schwartz, M. L.; Kajihara, H.; Norenburg, J. L. (2019). Molecular systematics of the heteronemertean genus
Dushia (Nemertea, Pilidiophora), with descriptions of
D. wijnhoffae sp. nov. and
D. nigra species complex comb. nov.
Zootaxa. 4691(4): 333-358.