Emberton, K.C. & Griffiths, O.L. (2009). Pulmonate land mollusks from three rainforest transects in northeastern Madagascar: part 1 of 2, with descriptions of 12 new species and one new subspecies: (Gastropoda: Pulmonata). Archiv für Molluskenkunde 138(2): 137–167.
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Emberton, K.C. & Griffiths, O.L.
2009
Pulmonate land mollusks from three rainforest transects in northeastern Madagascar: part 1 of 2, with descriptions of 12 new species and one new subspecies: (Gastropoda: Pulmonata).