World Register of marine Cave Species (WoRCS)

1. Species Spiroloculina abatholopa Loeblich & Tappan, 1994
2. Species Spiroloculina acescata Cushman, 1932
3. Species Spiroloculina aequa Cushman, 1932
4. Species Spiroloculina aequiformis McCulloch, 1977
5. Species Spiroloculina albatrossi Cushman & Todd, 1944
6. Species Spiroloculina anderseni Todd & Brönnimann, 1957
7. Species Spiroloculina angulariformis McCulloch, 1981
8. Species Spiroloculina angulata Cushman, 1917
9. Species Spiroloculina angulatiformis McCulloch, 1977
10. Species Spiroloculina angulosa Terquem, 1878
11. Species Spiroloculina angulosa d'Orbigny in Fornasini, 1904 (accepted > unreplaced junior homonym, Primary junior homonym of Spiroloculina angulosa Terquem, 1878)
12. Species Spiroloculina angusteoralis Parr, 1950
13. Species Spiroloculina antillarum d'Orbigny, 1839
14. Species Spiroloculina aperta Cushman & Todd, 1944
15. Species Spiroloculina atlantica Cushman, 1947
16. Species Spiroloculina barbariana McCulloch, 1977
17. Species Spiroloculina berthelotiana Wiesner, 1923
18. Species Spiroloculina biconcava McCulloch, 1977
19. Species Spiroloculina biformis Shchedrina, 1984
20. Species Spiroloculina boonei McCulloch, 1977
21. Species Spiroloculina bradyi Barker, 1960
22. Species Spiroloculina braithwaitensis McCulloch, 1977
23. Species Spiroloculina caduca Cushman, 1922
24. Species Spiroloculina caledoniana McCulloch, 1981
25. Species Spiroloculina californica Cushman & Todd, 1944
26. Species Spiroloculina canaliculata d'Orbigny, 1846
27. Species Spiroloculina carinata Fornasini, 1905
28. Species Spiroloculina caudata Ehrenberg, 1845
29. Species Spiroloculina charlesensis McCulloch, 1977
30. Species Spiroloculina chincaensis McCulloch, 1977
31. Species Spiroloculina clara Cushman, 1932
32. Species Spiroloculina colomi Said & Kamel, 1956
33. Species Spiroloculina communis Cushman & Todd, 1944
34. Species Spiroloculina concepcionensis McCulloch, 1977
35. Species Spiroloculina convexa Said, 1949
36. Species Spiroloculina corrugata Cushman & Todd, 1944
37. Species Spiroloculina cortesiana McCulloch, 1977
38. Species Spiroloculina cortesica McCulloch, 1977
39. Species Spiroloculina cristobalensis McCulloch, 1977
40. Species Spiroloculina curvata McCulloch, 1977
41. Species Spiroloculina cushmani Hada, 1931
42. Species Spiroloculina cymbium d'Orbigny, 1839
43. Species Spiroloculina dentata Cushman & Todd, 1944
44. Species Spiroloculina depressa d'Orbigny, 1826
45. Species Spiroloculina dilatata d'Orbigny, 1846
46. Species Spiroloculina dispar McCulloch, 1981
47. Species Spiroloculina dissidens McCulloch, 1977
48. Species Spiroloculina distincta McCulloch, 1977
49. Species Spiroloculina domantayi McCulloch, 1977
50. Species Spiroloculina ecuadorensis McCulloch, 1977
51. Species Spiroloculina elegantissima Said, 1949
52. Species Spiroloculina elevata Wiesner, 1923
53. Species Spiroloculina elongata d'Orbigny, 1852 (accepted > unreplaced junior homonym, Primary junior homonym of Spiroloculina elongata Ehrenberg, 1844)
54. Species Spiroloculina estebani Tinoco, 1958
55. Species Spiroloculina excavata d'Orbigny, 1846
56. Species Spiroloculina excisa Cushman & Todd, 1944
57. Species Spiroloculina eximia Cushman, 1922
58. Species Spiroloculina folium Todd, 1957
59. Species Spiroloculina foveolata Egger, 1893
60. Species Spiroloculina fragilis Uchio, 1960
61. Species Spiroloculina frailesiana McCulloch, 1977
62. Species Spiroloculina granulata Mikhalevich, 1976
63. Species Spiroloculina grateloupi d'Orbigny, 1852
64. Species Spiroloculina guppyi Todd & Brönnimann, 1957
65. Species Spiroloculina hadai Thalmann, 1933
66. Species Spiroloculina halmosae McCulloch, 1977
67. Species Spiroloculina hancocki McCulloch, 1981
68. Species Spiroloculina henbesti Petri, 1955
69. Species Spiroloculina hyalina Schulze, 1875
70. Species Spiroloculina ildefonsoensis McCulloch, 1977
71. Species Spiroloculina impressa Terquem, 1878
72. Species Spiroloculina incisa Cushman, 1921
73. Species Spiroloculina indica Cushman & Todd, 1944
74. Species Spiroloculina inflatiformis McCulloch, 1977
75. Species Spiroloculina insularis McCulloch, 1977
76. Species Spiroloculina jamesbayensis McCulloch, 1977
77. Species Spiroloculina jucunda Cushman & Ellisor, 1944
78. Species Spiroloculina krumbachi Wiesner, 1911
79. Species Spiroloculina lancea Ehrenberg, 1861
80. Species Spiroloculina latiformis McCulloch, 1977
81. Species Spiroloculina lavelaensis McCulloch, 1981
82. Species Spiroloculina libyca Martinotti, 1921
83. Species Spiroloculina limbata d'Orbigny, 1826
84. Species Spiroloculina lucida Cushman & Todd, 1944
85. Species Spiroloculina lyra d'Orbigny, 1852
86. Species Spiroloculina manifesta Cushman & Todd, 1944
87. Species Spiroloculina marshallana Todd, 1954
88. Species Spiroloculina mayori Cushman, 1924
89. Species Spiroloculina milletti Wiesner, 1912
90. Species Spiroloculina moresbyensis Rasheed, 1971
91. Species Spiroloculina mosesi Tinoco, 1958
92. Species Spiroloculina neocircularis McCulloch, 1977
93. Species Spiroloculina neoprofunda McCulloch, 1981
94. Species Spiroloculina nitida d'Orbigny, 1826
95. Species Spiroloculina norvegica Cushman & Todd, 1944
96. Species Spiroloculina novozealandica Cushman & Todd, 1944
97. Species Spiroloculina nuda Cushman, 1917
98. Species Spiroloculina nummiformis Said, 1949
99. Species Spiroloculina orbis Cushman, 1921
100. Species Spiroloculina ornata d'Orbigny, 1839
101. Species Spiroloculina ornatiformis McCulloch, 1977
102. Species Spiroloculina parvula Chapman, 1902
103. Species Spiroloculina pasquai Saidova, 1975
104. Species Spiroloculina pellucida Said, 1950
105. Species Spiroloculina penangensis McCulloch, 1977
106. Species Spiroloculina penangiana McCulloch, 1977
107. Species Spiroloculina pilata McCulloch, 1977
108. Species Spiroloculina pinasbayensis McCulloch, 1977
109. Species Spiroloculina planissima Wiesner, 1912
110. Species Spiroloculina planocommunis McCulloch, 1981
111. Species Spiroloculina planoconvexa Zheng, 1979
112. Species Spiroloculina planulata (Lamarck, 1804)
113. Species Spiroloculina plicata d'Orbigny, 1826
114. Species Spiroloculina poeyana d'Orbigny, 1839
115. Species Spiroloculina portparkerensis McCulloch, 1977
116. Species Spiroloculina pseudodepressa Mikhalevich, 1973
117. Species Spiroloculina pulchra Shchedrina, 1984
118. Species Spiroloculina regularis Cushman & Todd, 1944
119. Species Spiroloculina reticosa Chapman, 1915
120. Species Spiroloculina robusta Brady, 1884
121. Species Spiroloculina rostrata Reuss, 1850
122. Species Spiroloculina rotunda d'Orbigny, 1826
123. Species Spiroloculina rugosa Cushman & Todd, 1944
124. Species Spiroloculina samoaensis Cushman, 1924
125. Species Spiroloculina sanbenitoensis McCulloch, 1977
126. Species Spiroloculina sannicolasensis McCulloch, 1977
127. Species Spiroloculina scrobiculata Cushman, 1921
128. Species Spiroloculina serrulata Cushman, 1932
129. Species Spiroloculina singaporiana McCulloch, 1977
130. Species Spiroloculina soldanii Fornasini, 1886
131. Species Spiroloculina stabilis Zheng, 1988
132. Species Spiroloculina striata d'Orbigny in Fischer, 1870
133. Species Spiroloculina subaequa McCulloch, 1977
134. Species Spiroloculina subcommunis Mikhalevich, 1973
135. Species Spiroloculina subcommunis McCulloch, 1981 (accepted > unreplaced junior homonym, Primary junior homonym of Spiroloculina subcommunis Mikhalevich, 1973)
136. Species Spiroloculina subdepressa Mikhalevich, 1973
137. Species Spiroloculina subimpressa Parr, 1950
138. Species Spiroloculina sublimbata Parr, 1950
139. Species Spiroloculina tenuimargo Cushman, 1917
140. Species Spiroloculina tenuiseptata Brady, 1884
141. Species Spiroloculina terquemiana Fornasini, 1900
142. Species Spiroloculina tobagoensis McCulloch, 1981
143. Species Spiroloculina tortuosa Chapman, 1901
144. Species Spiroloculina utriaensis McCulloch, 1977
145. Species Spiroloculina valida Cushman, 1921
146. Species Spiroloculina venusta Cushman & Todd, 1944
147. Species Spiroloculina acutimargo Brady, 1884 accepted as Spirophthalmidium acutimargo (Brady, 1884)
148. Species Spiroloculina advena Cushman, 1921 accepted as Inaequalina advena (Cushman, 1921) (Opinion of Łuczkowska (1971))
149. Species Spiroloculina affixa Terquem, 1878 accepted as Inaequalina affixa (Terquem, 1878) (Opinion of Łuczkowska (1971))
150. Species Spiroloculina africana Martinotti, 1921 accepted as Spiroloculina soldanii Fornasini, 1886 (Subjective junior synonym Opinion of Cushman and Todd (1944))
151. Species Spiroloculina angulosa d'Orbigny, 1826 accepted as Spiroloculina angulosa d'Orbigny in Fornasini, 1904 (Nomen nudum)
152. Species Spiroloculina antillarium d'Orbigny, 1839 accepted as Spiroloculina antillarum d'Orbigny, 1839 (Incorrect subsequent spelling)
153. Species Spiroloculina arenaria Brady, 1884 accepted as Sigmoilopsis arenaria (Brady, 1884) (Opinion of Łuczkowska (1974), Łuczkowska in Jones (1994))
154. Species Spiroloculina arenata Cushman, 1921 accepted as Sigmoilopsis arenata (Cushman, 1921)
155. Species Spiroloculina asperula Karrer, 1868 accepted as Spirosigmoilina asperula (Karrer, 1868) accepted as Spiroglutina asperula (Karrer, 1868) (type species of Spiroglutina )
156. Species Spiroloculina attenuata Cushman & Todd, 1944 accepted as Naxotia attenuata (Cushman & Todd, 1944)
157. Species Spiroloculina barthelotiana Wiesner, 1923 accepted as Spiroloculina berthelotiana Wiesner, 1923 (Incorrect subsequent spelling)
158. Species Spiroloculina bohaiensis Zheng et al., 1978 accepted as Massilina penglaiensis (Jacot, 1925) (Opinion of Zheng & Fu, 2008)
159. Species Spiroloculina celata Costa, 1855 accepted as Sigmoilopsis celata (Costa, 1855)
160. Species Spiroloculina circularis Cushman, 1921 accepted as Spiroloculina regularis Cushman & Todd, 1944 (unaccepted > junior homonym, Nomen novum in Cushman and Todd (1944) Junior homonym of S. dorsata var. circularis Chapman, 1915 (already preoccupied by S. circularis Toula, 1914 [not figured]))
161. Species Spiroloculina concava Wiesner, 1916 accepted as Ophthalmidium concavum (Wiesner, 1916)
162. Species Spiroloculina concava Petri, 1954 accepted as Spiroloculina henbesti Petri, 1955 (unaccepted > junior homonym, New name in Thalman (1955) Junior homonym of S. acutimargo var. concava Wiesner, 1916)
163. Species Spiroloculina convexiuscula Brady, 1884 accepted as Nodobaculariella convexiuscula (Brady, 1884) (Opinion of Cushman and Todd (1944))
164. Species Spiroloculina costata Hada, 1931 accepted as Spiroloculina hadai Thalmann, 1933 (unaccepted > junior homonym, Nomen novum in Thalmann (1933) Junior homonym of S. costata Terquem, 1882)
165. Species Spiroloculina costifera Cushman, 1917 accepted as Ptychomiliola costifera (Cushman, 1917) (Opinion of Cushman and Todd (1944))
166. Species Spiroloculina crenata Karrer, 1868 accepted as Massilina crenata (Karrer, 1868) (Opinion of Rhumbler (1911))
167. Species Spiroloculina disparilis Terquem, 1878 accepted as Inaequalina disparilis (Terquem, 1878) (Opinion of Łuczkowska (1971))
168. Species Spiroloculina elegans Cushman, 1918 accepted as Spiroloculina foveolata Egger, 1893 (unaccepted > junior subjective synonym, Junior subjective synonym Opinion of Mamo (2016) [Primary junior homonym of Spiroloculina elegans Silvestri, 1896])
169. Species Spiroloculina elongata d'Orbigny, 1826 accepted as Spiroloculina elongata d'Orbigny, 1852 (Nomen nudum)
170. Species Spiroloculina folia Todd, 1957 accepted as Spiroloculina folium Todd, 1957 ('folium' neutral noun II declension)
171. Species Spiroloculina fragilissima Brady, 1884 accepted as Hauerina fragilissima (Brady, 1884) accepted as Parahauerinoides fragilissima (Brady, 1884) accepted as Pseudohauerina fragilissima (Brady, 1884) (Opinion of Cushman and Todd (1944))
172. Species Spiroloculina grata Terquem, 1878 accepted as Sigmoilina grata (Terquem, 1878)
173. Species Spiroloculina grateloupi d'Orbigny, 1826 accepted as Spiroloculina grateloupi d'Orbigny, 1852 (Nomen nudum)
174. Species Spiroloculina gratteloupi d'Orbigny, 1852 accepted as Spiroloculina grateloupi d'Orbigny, 1852 (Incorrect original spelling; species dedicated to Grateloup, Jean-Pierre-Sylvestre de (1782-1861) )
175. Species Spiroloculina henbesti Thalmann, 1955 accepted as Spiroloculina henbesti Petri, 1955 (New name in Thalman (1955) with the autorization of S. Petri who remains the author)
176. Species Spiroloculina inaequilateralis Schlumberger, 1893 accepted as Inaequalina inaequilateralis (Schlumberger, 1893) (Type species of inaequalina)
177. Species Spiroloculina laevigata Cushman & Todd, 1944 accepted as Massilina laevigata (Cushman & Todd, 1944) (Opinion of Zheng & Fu, 2008)
178. Species Spiroloculina obscura Cushman & Todd, 1944 accepted as Spiroloculina grateloupi d'Orbigny, 1852 (Invalid nomen novum. Cushman thought d'Orbigny's species was a junior homonym of Terquem's S. grateloupi (1878) not taking into account the description from 1852 (only Fornasini's 1904))
179. Species Spiroloculina penglaiensis Jacot, 1925 accepted as Massilina penglaiensis (Jacot, 1925) (Opinion of Zheng & Fu, 2008)
180. Species Spiroloculina poeyiana d'Orbigny, 1839 accepted as Spiroloculina poeyana d'Orbigny, 1839 (Incorrect original spelling)
181. Species Spiroloculina pusilla Earland, 1934 accepted as Spirosigmoilina pusilla (Earland, 1934) (Opinion of Jones (1994))
182. Species Spiroloculina rotundata Kreuzberg, 1930 accepted as Spiroloculina novozealandica Cushman & Todd, 1944 (unaccepted > junior homonym, Thought to be a junior homonym of a Williamson's subspecies (a misspelling) but a primary junior homonym of Spiroloculina excavata var. rotundata Costa, 1856)
183. Species Spiroloculina rotundata d'Orbigny, 1826 accepted as Spiroloculina rotunda d'Orbigny, 1826 (unaccepted > misspelling - incorrect subsequent spelling, rotundata is a misspelling of 'rotunda' in Williamson (1858))
184. Species Spiroloculina scita Cushman & Todd, 1944 accepted as Spiroloculina striata d'Orbigny in Fischer, 1870 (Invalid nomen novum. Cushman and Todd overlooked that S. striata d'Orbigny was a valid name in Fischer (1870) [not in Fornasini (1904)] before S. striata Terquem, 1882)
185. Species Spiroloculina speciosa Karrer, 1868 accepted as Spirosigmoilina speciosa (Karrer, 1868)
186. Species Spiroloculina striata d'Orbigny, 1826 accepted as Spiroloculina striata d'Orbigny in Fischer, 1870 (Nomen nudum)
187. Species Spiroloculina tenuis (Cžjžek, 1848) accepted as Spirosigmoilina tenuis (Cžjžek, 1848)
188. Species Spiroloculina tenuissima Reuss, 1867 accepted as Sigmoilinita tenuissima (Reuss, 1867) (Opinion of Łuczkowska (1974))
189. Species Spiroloculina terquemiana Heron-Allen & Earland, 1911 accepted as Spiroloculina carinata Le Calvez, 1947 † (Invalid nomen novum in Heron-Allen and Earland (1911) also preoccupied by S. terquemiana Fornasini, 1900)
190. Species Spiroloculina tripolitana Martinotti, 1921 accepted as Spiroloculina angulosa Terquem, 1878 (Subjective junior synonym Opinion of Cushman and Todd (1944))
191. Species Spiroloculina unicostata Cushman, 1917 accepted as Spirophthalmidium unicostatum (Cushman, 1917) (Opinion of Cushman and Todd (1944))
192. Species Spiroloculina validus Cushman, 1921 accepted as Spiroloculina valida Cushman, 1921 (Opinion of Walter (1960) Genus is feminine)
193. Species Spiroloculina ambullaris Ehrenberg, 1843 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
194. Species Spiroloculina ampla Terquem, 1880 (uncertain > nomen dubium, Opinion of Lévy et al. (1975))
195. Species Spiroloculina arenilega Rhumbler, 1949 (uncertain, Not described Invalid)
196. Species Spiroloculina complanata Egger, 1893 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
197. Species Spiroloculina flexuosa Ehrenberg, 1858 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
198. Species Spiroloculina galeata Ehrenberg, 1844 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
199. Species Spiroloculina interrupta Terquem, 1878 (uncertain > nomen dubium, Opinion of Mangin (1960) [broken type; nondiagnostic figure; teratological form] Subjective junior synonym of S. rotunda Opinion of Cushman and Todd (1944))
200. Species Spiroloculina involuta Egger, 1893 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
201. Species Spiroloculina lagena Ehrenberg, 1843 (uncertain > taxon inquirendum, Opinion of Cushman and Todd (1944) [Milioline stage of ?])
202. Species Spiroloculina lamella Egger, 1893 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
203. Species Spiroloculina laminata Terquem, 1880 (uncertain > nomen dubium, Opinion of Lévy et al. (1975))
204. Species Spiroloculina minor Ehrenberg, 1840 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
205. Species Spiroloculina orbicularis Ehrenberg, 1845 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))
206. Species Spiroloculina tuba Ehrenberg, 1844 (uncertain > nomen dubium, Opinion of Cushman and Todd (1944))

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