Canadian Register of Marine Species - Phyllophoridae Östergren, 1907

WoRMS taxon details

Phyllophoridae Östergren, 1907

AphiaID

123188 (urn:lsid:marinespecies.org:taxname:123188)

Classification

Status

accepted

Rank

Family

Typetaxon

Phyllophorus Grube, 1840

Parent

Dendrochirotida

All children (459)

Ordering

Alphabetically
By status

Children Display

direct main ranks all

Genus Allothyone Panning, 1949
1. Species Allothyone longicauda (Östergren, 1898)
2. Species Allothyone mexicana (Deichmann, 1946)
3. Species Allothyone mucronata (Sluiter, 1901)
4. Species Allothyone multipes (Augustin, 1908)
5. Species Allothyone presleri O'Loughlin in O'Loughlin, Stępień, Kuźniak & VandenSpiegel, 2013
6. Species Allothyone spadix (Sluiter, 1901)
Genus Anthochirus Chang, 1948
1. Species Anthochirus loui Chang, 1948
Genus Cladolella Heding & Panning, 1954
1. Species Cladolella virgo Heding & Panning, 1954
Genus Ekmanothyone Massin, 1993
1. Species Ekmanothyone incurva (Cherbonnier, 1988)
Genus Hemithyone Pawson, 1963
1. Species Hemithyone semperi (Bell, 1884)
Genus Lipotrapeza Clark, 1938
1. Species Lipotrapeza capilla Cherbonnier, 1958
2. Species Lipotrapeza eichleri O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
3. Species Lipotrapeza incurva Cherbonnier, 1988
4. Species Lipotrapeza japonica Heding & Panning, 1954
5. Species Lipotrapeza littoralis Yamana & Kohtsuka, 2017
6. Species Lipotrapeza litusi O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
7. Species Lipotrapeza purpurata Yamana & Kohtsuka, 2018
8. Species Lipotrapeza vestiens (Joshua, 1914)
9. Species Lipotrapeza ambigua Cherbonnier, 1988 accepted as Phyrella ambigua (Cherbonnier, 1988)
10. Species Lipotrapeza dearmatus (Dendy & Hindle, 1907) accepted as Neothyonidium dearmatum (Dendy & Hindle, 1907) (unaccepted > superseded combination)
11. Species Lipotrapeza ventripes (Joshua & Creed, 1915) accepted as Phyllostauros vercoi (Joshua & Creed, 1915)
Genus Massinium Samyn & Thandar, 2003
1. Species Massinium albicans Samyn, Thandar & VandenSpiegel, 2010
2. Species Massinium arthroprocessum (Thandar, 1989)
3. Species Massinium bonapartum O'Loughlin in O'Loughlin et al., 2014
4. Species Massinium dissimilis (Cherbonnier, 1988)
5. Species Massinium granulosum Samyn, Thandar & VandenSpiegel, 2010
6. Species Massinium japonicum Yamana, Ota, Kohtsuka & Yoshizaki, 2020
7. Species Massinium keesingi O'Loughlin in O'Loughlin et al., 2014
8. Species Massinium maculosum Samyn & Thandar, 2003
9. Species Massinium magnum (Ludwig, 1882)
10. Species Massinium melanieae O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
11. Species Massinium ocumichoensis Solís-Marín, Laguarda-Figueras, Conejeros-Vargas, Caballero-Ochoa & Durán-González, 2021
12. Species Massinium toyoshiomaruae Yamana, Ogawa & Ohtsuka in Yamana et al. 2021
13. Species Massinium vimsi O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
14. Species Massinium watsonae O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
Genus Neopentadactyla Deichmann, 1944
1. Species Neopentadactyla mixta (Östergren, 1898) Deichmann, 1944
Genus Neothyonidium Deichmann, 1938
1. Species Neothyonidium armatum Pawson, 1965
2. Species Neothyonidium dearmatum (Dendy & Hindle, 1907)
3. Species Neothyonidium hawaiiense (Fisher, 1907)
4. Species Neothyonidium inflatum (Sluiter, 1901)
5. Species Neothyonidium insolitum O'Loughlin in O'Loughlin et al., 2016
6. Species Neothyonidium intermedium (Koehler & Vaney, 1908)
7. Species Neothyonidium minutum (Ohshima, 1915)
8. Species Neothyonidium parvipedum Rowe, 1989
9. Species Neothyonidium parvum (Ludwig, 1881)
10. Species Neothyonidium spiniferum Liao & Pawson, 2001
11. Species Neothyonidium vultur (Sluiter, 1914)
12. Species Neothyonidium arthroprocessum Thandar, 1989 accepted as Massinium arthroprocessum (Thandar, 1989)
13. Species Neothyonidium dearmatus (Dendy & Hindle, 1907) accepted as Neothyonidium dearmatum (Dendy & Hindle, 1907) (misspelling)
14. Species Neothyonidium dissimilis Cherbonnier, 1988 accepted as Massinium dissimilis (Cherbonnier, 1988)
15. Species Neothyonidium magnum (Ludwig, 1882) accepted as Massinium magnum (Ludwig, 1882)
Genus Pentadactyla Hutton, 1878
1. Species Pentadactyla japonica (von Marenzeller, 1882)
2. Species Pentadactyla longidentis (Hutton, 1872)
Genus Pentamera Ayres, 1852
1. Species Pentamera beebei Deichmann, 1938
2. Species Pentamera calcigera (Stimpson, 1851)
3. Species Pentamera charlottae Deichmann, 1938
4. Species Pentamera chierchiae (Ludwig, 1886)
5. Species Pentamera chiloensis (Ludwig, 1886)
6. Species Pentamera citrea (Semper, 1867)
7. Species Pentamera constricta (Ohshima, 1915)
8. Species Pentamera dubia Cherbonnier, 1951
9. Species Pentamera fonsecae Solis-Marin, Alvarado, Conejeros-Vargas & Caballero-Ochoa, 2020
10. Species Pentamera lissoplaca (Clark, 1924)
11. Species Pentamera misakiensis Yamana & Kohtsuka, 2018
12. Species Pentamera montereyensis Deichmann, 1938
13. Species Pentamera obscura Cherbonnier, 1951
14. Species Pentamera paraibanensis Prata & Christofferson, 2016
15. Species Pentamera pediparva Lambert, 1998
16. Species Pentamera populifera (Stimpson, 1864)
17. Species Pentamera pseudocalcigera Deichmann, 1938
18. Species Pentamera pseudopopulifera Deichmann, 1938
19. Species Pentamera pulcherrima Ayres, 1852
20. Species Pentamera rigida Lambert, 1998
21. Species Pentamera trachyplaca (Clark, 1924)
22. Species Pentamera zacae Deichmann, 1938
23. Species Pentamera chierchia (Ludwig, 1887) accepted as Pentamera chierchiae (Ludwig, 1886) (misspelling)
24. Species Pentamera chuni (Ludwig & Heding, 1935) accepted as Temparena chuni (Ludwig & Heding, 1935)
25. Species Pentamera velligera (Ludwig & Heding, 1935) accepted as Sclerothyone velligera (Ludwig & Heding, 1935)
Genus Phyllophorella Heding & Panning, 1954
1. Species Phyllophorella drachi (Cherbonnier & Guille, 1968)
2. Species Phyllophorella dubius (Cherbonnier, 1961)
3. Species Phyllophorella kohkutiensis (Heding & Panning, 1954)
4. Species Phyllophorella liuwutiensis (Yang, 1937)
5. Species Phyllophorella longipeda (Semper, 1867)
6. Species Phyllophorella notialis O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
7. Species Phyllophorella perforata (Clark, 1932)
8. Species Phyllophorella purpureopunctata (Sluiter, 1901)
9. Species Phyllophorella robusta (Heding & Panning, 1954)
10. Species Phyllophorella rosetta (Thandar, 1994)
11. Species Phyllophorella roseus (Cherbonnier & Féral, 1981)
12. Species Phyllophorella spiculata (Chang, 1935)
13. Species Phyllophorella thyonoides (H.L. Clark, 1938) accepted as Phyrella thyonoides (H.L. Clark, 1938)
Subfamily Phyllophorinae Östergren, 1907
Genus Phyllophorus Grube, 1840
1. Subgenus Phyllophorus (Isophyllophorus) Liao & Pawson, 2001
  1. Species Phyllophorus (Isophyllophorus) orientalis Liao & Pawson, 2001
2. Subgenus Phyllophorus (Phyllonovus) Cherbonnier, 1988
  1. Species Phyllophorus (Phyllonovus) anomalia Cherbonnier, 1988
3. Subgenus Phyllophorus (Phyllophorus) Heding & Panning, 1954
  1. Species Phyllophorus (Phyllophorus) calypsoi Cherbonnier, 1954
  2. Species Phyllophorus (Phyllophorus) granulatus (Grube, 1840)
  3. Species Phyllophorus (Phyllophorus) maculatus Liao, Pawson & Liu, 2007
  4. Species Phyllophorus (Phyllophorus) pedinaequalis Cherbonnier, 1969
  5. Species Phyllophorus (Phyllophorus) urna Grube, 1840
  6. Species Phyllophorus (Phyllophorus) bedoti Koehler, 1895 accepted as Phyrella bedoti (Koehler, 1895)
  7. Species Phyllophorus (Phyllophorus) roseus Cherbonnier & Féral, 1981 accepted as Phyllophorella roseus (Cherbonnier & Féral, 1981)
4. Subgenus Phyllophorus (Phyllothuria) Heding & Panning, 1954
  1. Species Phyllophorus (Phyllothuria) cebuensis (Semper, 1867)
  2. Species Phyllophorus (Phyllothuria) celer koehler & Vaney, 1908
  3. Species Phyllophorus (Phyllothuria) discrepans (Sluiter, 1901)
  4. Species Phyllophorus (Phyllothuria) donghaiensis Liao & Pawson, 2001
  5. Species Phyllophorus (Phyllothuria) hypsipyrga (von Marenzeller, 1882)
  6. Species Phyllophorus (Phyllothuria) mammulus Cherbonnier, 1965
  7. Species Phyllophorus (Phyllothuria) ordinata Chang, 1935
5. Species Phyllophorus brocki Ludwig, 1888
6. Species Phyllophorus dubia (Bedford, 1899)
7. Species Phyllophorus exigua (Cherbonnier, 1988)
8. Species Phyllophorus holothurioides Ludwig, 1875
9. Species Phyllophorus kungi (O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012)
10. Species Phyllophorus proteus Bell, 1884
11. Species Phyllophorus tenuis Haacke, 1880
12. Subgenus Phyllophorus (Urodemella) Deichmann, 1944 (not available: no type species designated and post-1930 (ICZN 13.3))
  1. Species Phyllophorus (Urodemella) arenicola Pawson & Miller, 1992 accepted as Euthyonidiella arenicola (Pawson & Miller, 1992)
  2. Species Phyllophorus (Urodemella) brocki Ludwig, 1888 accepted as Phyllophorus brocki Ludwig, 1888 (Urodemella is not an available name)
  3. Species Phyllophorus (Urodemella) holothurioides Ludwig, 1875 accepted as Phyllophorus holothurioides Ludwig, 1875 (Urodemella is not an available name)
  4. Species Phyllophorus (Urodemella) occidentalis (Ludwig, 1875) accepted as Euthyonidiella occidentalis (Ludwig, 1875)
  5. Species Phyllophorus (Urodemella) oculus Cherbonnier, 1988 accepted as Ohshimella ocula (Cherbonnier, 1988)
  6. Species Phyllophorus (Urodemella) proteus Bell, 1884 accepted as Phyllophorus proteus Bell, 1884 (Urodemella is not an available name)
  7. Species Phyllophorus (Urodemella) tenuis Haacke, 1880 accepted as Phyllophorus tenuis Haacke, 1880 (Urodemella is not an available name)
13. Species Phyllophorus aculeatus Ludwig, 1894 accepted as Phyrella aculeata (Ludwig, 1894)
14. Species Phyllophorus ambiguus Heding, 1942 accepted as Euthyonidiella ambigua (Heding, 1942)
15. Species Phyllophorus aparecidae Ancona Lopez, 1962 accepted as Thyonidium seguroensis (Deichmann, 1930)
16. Species Phyllophorus communis (Forbes & Goodsir in Forbes, 1841) accepted as Cucumaria communis Forbes & Goodsir in Forbes, 1841
17. Species Phyllophorus contractura Cherbonnier, 1988 accepted as Phyrella ambigua (Cherbonnier, 1988)
18. Species Phyllophorus cornus Heding, 1934 accepted as Actinocucumis cornus (Heding, 1934)
19. Species Phyllophorus cylindricus Ohshima, 1915 accepted as Ekmania cylindricum (Ohshima, 1915)
20. Species Phyllophorus dearmatus Dendy & Hindle, 1907 accepted as Neothyonidium dearmatum (Dendy & Hindle, 1907)
21. Species Phyllophorus destichadus Deichmann, 1930 accepted as Euthyonidiella destichada (Deichmann, 1930)
22. Species Phyllophorus diomedeae Ohshima, 1915 accepted as Thyonidium diomedeae (Ohshima, 1915)
23. Species Phyllophorus dobsoni Bell, 1883 accepted as Globosita dobsoni (Bell, 1883)
24. Species Phyllophorus ehrenbergi Selenka, 1868 accepted as Ohshimella ehrenbergii (Selenka, 1868) (unaccepted > misspelling - incorrect subsequent spelling)
25. Species Phyllophorus fragilis Mitsukuri & Ohshima in Ohshima, 1912 accepted as Phyrella fragilis (Mitsukuri & Ohshima in Ohshima, 1912)
26. Species Phyllophorus frauenfeldi Ludwig, 1875 accepted as Ohshimella ehrenbergii (Selenka, 1868)
27. Species Phyllophorus glaucus Ohshima, 1915 accepted as Ekmania glaucum (Ohshima, 1915)
28. Species Phyllophorus granulatus (Grube, 1840) accepted as Phyllophorus (Phyllophorus) granulatus (Grube, 1840)
29. Species Phyllophorus hamatus Sluiter, 1914 accepted as Cladolabes hamatus (Sluiter, 1914)
30. Species Phyllophorus incompertus Théel, 1886 accepted as Cladolabes perspicillum (Selenka, 1867)
31. Species Phyllophorus inflatus Sluiter, 1901 accepted as Neothyonidium inflatum (Sluiter, 1901)
32. Species Phyllophorus intermedius Koehler & Vaney, 1908 accepted as Neothyonidium intermedium (Koehler & Vaney, 1908)
33. Species Phyllophorus limaconotus (Brandt, 1835) accepted as Cladolabes limaconotus Brandt, 1835 (unaccepted > superseded combination)
34. Species Phyllophorus mammulus Cherbonnier, 1965 accepted as Phyllophorus (Phyllothuria) mammulus Cherbonnier, 1965
35. Species Phyllophorus minutus Ohshima, 1915 accepted as Neothyonidium minutum (Ohshima, 1915)
36. Species Phyllophorus ordinatus Chang, 1935 accepted as Phyllophorus (Phyllothuria) ordinata Chang, 1935
37. Species Phyllophorus palmae Ancona Lopez, 1962 accepted as Parathyone surinamensis (Semper, 1867)
38. Species Phyllophorus panningi Domantay, 1953 (unaccepted > nomen nudum)
39. Species Phyllophorus parvipedes Clark, 1938 accepted as Phyllophorus (Phyllophorella) spiculata Chang, 1935
40. Species Phyllophorus seguroensis Deichmann, 1930 accepted as Thyonidium seguroensis (Deichmann, 1930)
41. Species Phyllophorus simplex Sluiter, 1914 accepted as Actinocucumis simplex (Sluiter, 1914)
42. Species Phyllophorus spiculatus Chang, 1935 accepted as Phyllophorus (Phyllophorella) spiculata Chang, 1935
43. Species Phyllophorus thyonoides H.L. Clark, 1938 accepted as Phyrella thyonoides (H.L. Clark, 1938)
44. Species Phyllophorus transvectus Sluiter, 1914 accepted as Afrocucumis africana (Semper, 1867)
45. Species Phyllophorus trapezus H.L. Clark, 1932 accepted as Phyrella trapeza (H.L. Clark, 1932)
46. Species Phyllophorus tungshanensis Yang, 1937 accepted as Euthyonidiella tungshanensis (Yang, 1937)
47. Species Phyllophorus ventripes Joshua & Creed, 1915 accepted as Phyllostauros vercoi (Joshua & Creed, 1915)
48. Species Phyllophorus vestiens Joshua, 1914 accepted as Lipotrapeza vestiens (Joshua, 1914)
49. Species Phyllophorus vultur Sluiter, 1914 accepted as Neothyonidium vultur (Sluiter, 1914)
50. Species Phyllophorus zacae Deichmann, 1938 accepted as Euthyonidiella zacae (Deichmann, 1938)
51. Subgenus Phyllophorus (Phyllophorella) Heding & Panning, 1954 represented as Phyllophorella Heding & Panning, 1954
  1. Species Phyllophorus (Phyllophorella) contractura Cherbonnier, 1988 accepted as Phyrella ambigua (Cherbonnier, 1988)
  2. Species Phyllophorus (Phyllophorella) kohkutiensis Heding & Panning, 1954 accepted as Phyllophorella kohkutiensis (Heding & Panning, 1954)
  3. Species Phyllophorus (Phyllophorella) liuwutiensis Yang, 1937 accepted as Phyllophorella liuwutiensis (Yang, 1937)
  4. Species Phyllophorus (Phyllophorella) roseus Cherbonnier & Féral, 1981 accepted as Phyllophorella roseus (Cherbonnier & Féral, 1981)
  5. Species Phyllophorus (Phyllophorella) drachi Cherbonnier & Guille, 1968 represented as Phyllophorella drachi (Cherbonnier & Guille, 1968)
  6. Species Phyllophorus (Phyllophorella) dubius Cherbonnier, 1961 represented as Phyllophorella dubius (Cherbonnier, 1961)
  7. Species Phyllophorus (Phyllophorella) longipeda (Semper, 1867) represented as Phyllophorella longipeda (Semper, 1867)
  8. Species Phyllophorus (Phyllophorella) perforata (H.L. Clark, 1932) represented as Phyllophorella perforata (Clark, 1932)
  9. Species Phyllophorus (Phyllophorella) purpureopunctata (Sluiter, 1901) represented as Phyllophorella purpureopunctata (Sluiter, 1901)
  10. Species Phyllophorus (Phyllophorella) robusta Heding & Panning, 1954 represented as Phyllophorella robusta (Heding & Panning, 1954)
  11. Species Phyllophorus (Phyllophorella) rosetta Thandar, 1994 represented as Phyllophorella rosetta (Thandar, 1994)
  12. Species Phyllophorus (Phyllophorella) spiculata Chang, 1935 represented as Phyllophorella spiculata (Chang, 1935)
52. Species Phyllophorus anomalia Cherbonnier, 1988 represented as Phyllophorus (Phyllonovus) anomalia Cherbonnier, 1988
53. Species Phyllophorus bedoti Koehler, 1895 represented as Phyllophorus (Phyllophorus) bedoti Koehler, 1895
54. Species Phyllophorus calypsoi Cherbonnier, 1954 represented as Phyllophorus (Phyllophorus) calypsoi Cherbonnier, 1954
55. Species Phyllophorus celer Koehler & Vaney, 1908 represented as Phyllophorus (Phyllothuria) celer koehler & Vaney, 1908
56. Species Phyllophorus drachi Cherbonnier & Guille, 1968 represented as Phyllophorus (Phyllophorella) drachi Cherbonnier & Guille, 1968
57. Species Phyllophorus kohkutiensis Heding & Panning, 1954 represented as Phyllophorus (Phyllophorella) kohkutiensis Heding & Panning, 1954
58. Species Phyllophorus liuwutiensis Yang, 1937 represented as Phyllophorus (Phyllophorella) liuwutiensis Yang, 1937
59. Species Phyllophorus occidentalis (Ludwig, 1875) represented as Phyllophorus (Urodemella) occidentalis (Ludwig, 1875)
60. Species Phyllophorus oculus Cherbonnier, 1988 represented as Phyllophorus (Urodemella) oculus Cherbonnier, 1988
61. Species Phyllophorus pedinaequalis Cherbonnier, 1969 represented as Phyllophorus (Phyllophorus) pedinaequalis Cherbonnier, 1969
62. Species Phyllophorus rosetta Thandar, 1994 represented as Phyllophorus (Phyllophorella) rosetta Thandar, 1994
63. Species Phyllophorus roseus Cherbonnier & Féral, 1981 represented as Phyllophorus (Phyllophorella) roseus Cherbonnier & Féral, 1981
64. Species Phyllophorus urna Grube, 1840 represented as Phyllophorus (Phyllophorus) urna Grube, 1840
65. Species Phyllophorus panningi Domantay, 1961 (uncertain > nomen dubium)
Genus Phyllostauros O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
1. Species Phyllostauros vercoi (Joshua & Creed, 1915)
Genus Phyrella Heding & Panning, 1954
1. Species Phyrella ambigua (Cherbonnier, 1988)
2. Species Phyrella bedoti (Koehler, 1895)
3. Species Phyrella drozdovi (Levin & Stepanov, 1999)
4. Species Phyrella fragilis (Mitsukuri & Ohshima in Ohshima, 1912)
5. Species Phyrella mookiei Michonneau & Paulay, 2014
6. Species Phyrella thyonoides (H.L. Clark, 1938)
7. Species Phyrella trapeza (H.L. Clark, 1932)
8. Species Phyrella aculeata (Ludwig, 1894) accepted as Euthyonidiella aculeata (Ludwig, 1894)
9. Species Phyrella tenera (Ludwig, 1875) (uncertain > taxon inquirendum)
Genus Pseudoplacothuria Yamana & Kohtsuka, 2018
1. Species Pseudoplacothuria ohshimai (Liao, 1997)
Genus Selenkiella Heding & Panning, 1954
1. Species Selenkiella malayense Heding & Panning, 1954
2. Species Selenkiella paradoxa Cherbonnier, 1970
3. Species Selenkiella siamense Heding & Panning, 1954
Subfamily Semperiellinae Heding & Panning, 1954
Genus Stolus Selenka, 1867
1. Species Stolus albescens Liao in Liao & Clark, 1995
2. Species Stolus axiologa (H.L. Clark, 1938)
3. Species Stolus buccalis (Stimpson, 1855)
4. Species Stolus canescens (Semper, 1867)
5. Species Stolus cognatus (Lampert, 1885)
6. Species Stolus conjungens (Semper, 1867)
7. Species Stolus crassus Liao & Pawson, 2001
8. Species Stolus dentatus Thandar, 2005
9. Species Stolus micronodosus Liao & Pawson, 2001
10. Species Stolus minutus (H.L. Clark, 1938)
11. Species Stolus papillatus (Sluiter, 1887)
12. Species Stolus parafusus (Deichmann, 1941)
13. Species Stolus pseudoalbescens Thandar, 2005
14. Species Stolus pseudofusus (Deichmann, 1930)
15. Species Stolus punctatus (Ohshima, 1915)
16. Species Stolus uniannulatus (Sluiter, 1914)
17. Species Stolus conjugens (Semper, 1868) accepted as Stolus conjungens (Semper, 1867) (misspelling)
18. Species Stolus firmus Selenka, 1867 accepted as Plesiocolochirus spinosus (Quoy & Gaimard, 1834)
19. Species Stolus gibber Selenka, 1867 accepted as Neothyone gibber (Selenka, 1867)
20. Species Stolus kilberti Rajpal & Thandar, 1999 accepted as Pseudostolus kilberti (Rajpal & Thandar, 1999)
21. Species Stolus ovulum Selenka, 1867 accepted as Afrocucumis ovulum (Selenka, 1867)
22. Species Stolus rapax (Koehler & Vaney, 1908) accepted as Pseudostolus rapax (Koehler & Vaney, 1908)
23. Species Stolus sacellus Selenka, 1867 accepted as Stolus buccalis (Stimpson, 1855)
Genus Thorsonia Heding, 1940
1. Species Thorsonia adversaria (Semper, 1867)
2. Species Thorsonia fusiformis Heding, 1940
3. Species Thorsonia investigatoris (Koehler & Vaney, 1908)
Genus Thyone Oken, 1815
1. Subgenus Thyone (Sclerodactyla)
  1. Species Thyone (Sclerodactyla) braziliensis Verrill, 1868 accepted as Ocnus braziliensis (Verrill, 1868)
2. Subgenus Thyone (Stolus)
  1. Species Thyone (Stolus) chilensis Semper, 1868 accepted as Eucyclus chilensis (Semper, 1868)
  2. Species Thyone (Stolus) mirabilis Ludwig, 1875 accepted as Havelockia versicolor (Semper, 1867)
  3. Species Thyone (Stolus) rigida Semper, 1867 accepted as Stolus buccalis (Stimpson, 1855)
  4. Species Thyone (Stolus) rosacea Semper, 1869 accepted as Staurothyone rosacea (Semper, 1869)
3. Subgenus Thyone (Thyone)
  1. Species Thyone (Thyone) suspecta Ludwig, 1875 accepted as Parathyone suspecta (Ludwig, 1875)
4. Subgenus Thyone (Thyonidium)
  1. Species Thyone (Thyonidium) Lechleri Lampert, 1885 accepted as Trachythyone lechleri (Lampert, 1885)
5. Species Thyone adinopoda Pawson & Miller, 1981
6. Species Thyone andrewsii (Farran, 1860)
7. Species Thyone anomala Östergren, 1898
8. Species Thyone aurea (Quoy & Gaimard, 1834)
9. Species Thyone avenusta Cherbonnier, 1970
10. Species Thyone bacescoi Cherbonnier, 1972
11. Species Thyone benti Deichmann, 1937
  1. Variety Thyone benti var. zacae Deichmann, 1938 accepted as Thyone benti Deichmann, 1937
12. Species Thyone bicornis Ohshima, 1915
13. Species Thyone bidentata Deichmann, 1941
14. Species Thyone brasiliana Prata, Manso & Christoffersen, 2020
15. Species Thyone carens Cherbonnier, 1988
16. Species Thyone cherbonnieri Reys, 1959
17. Species Thyone comata Cherbonnier, 1988
18. Species Thyone crassidisca Pawson & Miller, 1981
19. Species Thyone crebrapodia Cherbonnier, 1988
20. Species Thyone curvata Lampert, 1885
21. Species Thyone deichmannae Madsen, 1941
22. Species Thyone discolor Sluiter, 1901
23. Species Thyone dura Koehler & Vaney, 1908
24. Species Thyone flindersi O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
25. Species Thyone florianoi Martins & Tavares, 2018
26. Species Thyone fusca Pearson, 1903
27. Species Thyone fusus (O.F. Müller, 1776)
  1. Subspecies Thyone fusus chinensis Yang, 1937
  2. Subspecies Thyone fusus mediterranea Madsen, 1941
  3. Variety Thyone fusus var. chinensis Yang, 1937 accepted as Thyone fusus chinensis Yang, 1937
  4. Variety Thyone fusus var. mediterranea Madsen, 1941 accepted as Thyone fusus mediterranea Madsen, 1941
  5. Variety Thyone fusus var. papuensis Théel, 1886 accepted as Thyone papuensis Théel, 1886
28. Species Thyone gadeana Perrier R., 1898
29. Species Thyone grisea Clark, 1938
30. Species Thyone guillei Cherbonnier, 1988
31. Species Thyone herberti Thandar & Rajpal, 1999
32. Species Thyone hirta Cherbonnier, 1970
33. Species Thyone imperfecta (Cherbonnier, 1970)
34. Species Thyone inermis Heller, 1868
  1. Variety Thyone inermis var. coronensis Domantay, 1953 (unaccepted > nomen nudum)
  2. Variety Thyone inermis var. coronensis Domantay, 1961 (uncertain > nomen dubium)
35. Species Thyone infusca Cherbonnier, 1954
36. Species Thyone joshuai O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
37. Species Thyone kerkosa O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
38. Species Thyone kyusyuensis Yamana, Ogawa & Ohtsuka in Yamana et al. 2021
39. Species Thyone liaoi Yamana, Ogawa & Ohtsuka in Yamana et al. 2021
40. Species Thyone longicornis Cherbonnier, 1988
41. Species Thyone micra Clark, 1938
42. Species Thyone montoucheti Tommasi, 1971
43. Species Thyone neofusus Deichmann, 1941
44. Species Thyone nigra Joshua & Creed, 1915
45. Species Thyone okeni Bell, 1884
46. Species Thyone papuensis Théel, 1886
47. Species Thyone pawsoni Tommasi, 1972
48. Species Thyone pedata Semper, 1867
49. Species Thyone pohaiensis Liao, 1986
50. Species Thyone polybranchia Östergren, 1898
51. Species Thyone profusus Cherbonnier & Féral, 1981
52. Species Thyone propinqua Cherbonnier, 1970
53. Species Thyone purpureopunctata Liao & Pawson, 2001
54. Species Thyone quadruperforata Cherbonnier, 1954
55. Species Thyone roscovita Hérouard, 1889
56. Species Thyone sinensis Liao & Pawson, 2001
57. Species Thyone sineturra Cherbonnier, 1988
58. Species Thyone spenceri O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
59. Species Thyone spinifera Liao in Liao & Clark, 1995
60. Species Thyone strangeri Deichmann, 1941
61. Species Thyone susamiensis Yamana, Hirai & Hirashima, 2015
62. Species Thyone tanyspiera Pawson & Miller, 1988
63. Species Thyone theeli Rowe in Rowe & Gates, 1995
64. Species Thyone tourvillei O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012
65. Species Thyone toyoshiomaruae Yamana, Ogawa & Ohtsuka in Yamana et al. 2021
66. Species Thyone vadosa Cherbonnier, 1988
67. Species Thyone venusta Selenka, 1868
68. Species Thyone venustella Ludwig & Heding, 1935
69. Species Thyone villosa Semper, 1867
70. Species Thyone vitrea Sluiter, 1901
71. Species Thyone waltinhoi Martins & Souto, 2018
72. Species Thyone alba Clark, 1938 accepted as Thyone dura Koehler & Vaney, 1908
73. Species Thyone anomula Massin, 1987 accepted as Thyone anomala Östergren, 1898 (unaccepted > unavailable name, misspelling and incorrect attribution by Thandar & Rajpal 1999 of T. anomala Ostergren discussed in Massin 1987)
74. Species Thyone articulata Vaney, 1908 accepted as Thyonina articulata (Vaney, 1908)
75. Species Thyone axiologa Clark, 1938 accepted as Stolus axiologa (H.L. Clark, 1938)
76. Species Thyone belli Ludwig, 1886 accepted as Pseudothyone belli (Ludwig, 1886)
77. Species Thyone brevidentis Hutton, 1872 accepted as Squamocnus brevidentis (Hutton, 1872)
78. Species Thyone briareus (Lesueur, 1824) accepted as Holothuria briareus Lesueur, 1824
79. Species Thyone buccalis Stimpson, 1855 accepted as Stolus buccalis (Stimpson, 1855)
  1. Subspecies Thyone buccalis bourdesae Domantay, 1962 accepted as Stolus buccalis (Stimpson, 1855)
  2. Variety Thyone buccalis var. pallida Clark, 1938 accepted as Stolus buccalis (Stimpson, 1855)
80. Species Thyone calcarea Pearson, 1903 accepted as Havelockia versicolor (Semper, 1867)
81. Species Thyone castanea Lampert, 1889 accepted as Thyone papuensis Théel, 1886
82. Species Thyone caudata Hutton, 1872 accepted as Pentadactyla longidentis (Hutton, 1872)
83. Species Thyone ceciliae Domantay, 1953 (unaccepted > nomen nudum)
84. Species Thyone clarki Domantay, 1953 (unaccepted > nomen nudum)
85. Species Thyone cognata (Lampert, 1885) accepted as Stolus cognatus (Lampert, 1885) (superseded combination)
86. Species Thyone colnettensis Domantay, 1953 (unaccepted > nomen nudum)
87. Species Thyone constituta Sluiter, 1910 accepted as Phyllophorus (Urodemella) occidentalis (Ludwig, 1875)
88. Species Thyone cunninghami Bell, 1883 accepted as Hemioedema spectabilis (Ludwig, 1883)
89. Species Thyone elegans Norman, 1869 accepted as Havelockia inermis (Heller, 1868)
90. Species Thyone festina Koehler & Vaney, 1908 accepted as Havelockia festina (Koehler & Vaney, 1908)
91. Species Thyone flexus Hodge, 1865 accepted as Thyone fusus (O.F. Müller, 1776)
92. Species Thyone floccosa Norman, 1864 (unaccepted > nomen nudum)
93. Species Thyone floridana Domantay, 1953 (unaccepted > nomen nudum)
94. Species Thyone glasselli Deichmann, 1936 accepted as Athyone glasselli (Deichmann, 1936)
95. Species Thyone hassleri Théel, 1886 accepted as Trachythyone lechleri (Lampert, 1885)
96. Species Thyone hornelli Pearson, 1903 accepted as Phyllophorus (Urodemella) brocki Ludwig, 1888
97. Species Thyone imbricata Ohshima, 1915 accepted as Stereoderma imbricata (Ohshima, 1915)
98. Species Thyone longidentis Hutton, 1872 accepted as Pentadactyla longidentis (Hutton, 1872)
99. Species Thyone lugubris Deichmann, 1939 accepted as Pachythyone lugubris (Deichmann, 1939)
100. Species Thyone meridionalis Bell, 1883 accepted as Hemioedema spectabilis (Ludwig, 1883)
101. Species Thyone mexicana Deichmann, 1946 accepted as Allothyone mexicana (Deichmann, 1946)
102. Species Thyone micropunctata Sluiter, 1910 accepted as Pseudothyone belli (Ludwig, 1886)
103. Species Thyone minuta H.L. Clark, 1938 accepted as Stolus minutus (H.L. Clark, 1938)
104. Species Thyone montereyensis (Deichmann, 1938) accepted as Pentamera montereyensis Deichmann, 1938
105. Species Thyone multipes Augustin, 1908 accepted as Allothyone multipes (Augustin, 1908)
106. Species Thyone muricata Lampert, 1886 [non Studer, 1876] accepted as Heterocucumis steineni (Ludwig, 1898) (Misidentification)
107. Species Thyone panamensis Ludwig, 1886 accepted as Neothyone panamensis (Ludwig, 1886)
108. Species Thyone papillata Sluiter, 1887 accepted as Stolus papillatus (Sluiter, 1887)
109. Species Thyone parafusus Deichmann, 1941 accepted as Stolus parafusus (Deichmann, 1941)
110. Species Thyone parva Ohshima, 1915 accepted as Paracucumaria parva (Ohshima, 1915)
111. Species Thyone perforata Clark, 1932 accepted as Phyllophorus (Phyllophorella) perforata (H.L. Clark, 1932)
112. Species Thyone perissa Clark, 1938 accepted as Massinium magnum (Ludwig, 1882)
113. Species Thyone pervicax Théel, 1886 accepted as Thyonella pervicax (Théel, 1886)
114. Species Thyone pituitosa Sluiter, 1901 accepted as Havelockia pituitosa (Sluiter, 1901)
115. Species Thyone portlockii Forbes, 1841 accepted as Thyonidium drummondii (Thompson, 1840) (synonym)
116. Species Thyone poucheti Barrois, 1882 accepted as Pseudothyone raphanus (Düben & Koren, 1846) (synonym)
117. Species Thyone proceracorona Cherbonnier, 1952 accepted as Thyone aurea (Quoy & Gaimard, 1834)
118. Species Thyone pseudofusus Deichmann, 1930 accepted as Stolus pseudofusus (Deichmann, 1930)
119. Species Thyone punctata Ohshima, 1915 accepted as Stolus punctatus (Ohshima, 1915)
120. Species Thyone raphanus Düben & Koren, 1846 accepted as Pseudothyone raphanus (Düben & Koren, 1846) (synonym)
121. Species Thyone recurvata Théel, 1886 accepted as Trachythyone muricata Studer, 1876 (junior synonym)
122. Species Thyone rubra Clark, 1901 accepted as Pachythyone rubra (Clark, 1901)
123. Species Thyone sabanillaensis Deichmann, 1930 accepted as Thyonella sabanillaensis (Deichmann, 1930)
124. Species Thyone sacellus (Selenka) accepted as Thyone buccalis Stimpson, 1855
125. Species Thyone sargassi Lampert, 1889 accepted as Staurothyone rosacea (Semper, 1869)
126. Species Thyone scabra Verrill, 1873 accepted as Havelockia scabra (Verrill, 1873)
127. Species Thyone scotiae Vaney, 1906 accepted as Crucella scotiae (Vaney, 1906)
128. Species Thyone serratus Britten, 1910 accepted as Thyone aurea (Quoy & Gaimard, 1834)
129. Species Thyone serrifera Östergren, 1898 accepted as Pseudothyone serrifera (Östergren, 1898) (synonym)
130. Species Thyone similis Ludwig, 1886 accepted as Neothyone gibber (Selenka, 1867)
131. Species Thyone sluiteri Lampert, 1889 accepted as Phyllophorus (Urodemella) proteus Bell, 1884
132. Species Thyone solida Deichmann, 1930 accepted as Euthyonacta solida (Deichmann, 1930)
133. Species Thyone spadix Sluiter, 1901 accepted as Allothyone spadix (Sluiter, 1901)
134. Species Thyone spectabilis Ludwig, 1883 accepted as Hemioedema spectabilis (Ludwig, 1883)
135. Species Thyone subvillosa Hérouard, 1889 accepted as Thyone fusus (O.F. Müller, 1776) (synonym)
136. Species Thyone surinamensis Semper, 1867 accepted as Parathyone surinamensis (Semper, 1867)
137. Species Thyone tanyspeira Pawson, 1976 accepted as Thyone tanyspiera Pawson & Miller, 1988 (misspelling and misattribution)
138. Species Thyone tenella Selenka, 1867 accepted as Sclerodactyla briareus (Lesueur, 1824)
139. Species Thyone trita Sluiter, 1910 accepted as Euthyonidiella trita (Sluiter, 1910)
140. Species Thyone turricata Vaney, 1906 accepted as Paracucumis turricata (Vaney, 1906)
  1. Variety Thyone turricata var. lourdesae Domantay, 1953 (unaccepted > nomen nudum)
  2. Variety Thyone turricata var. lourdesae Domantay, 1961 (uncertain > nomen dubium)
141. Species Thyone turrissolida Cherbonnier, 1954 accepted as Thyone aurea (Quoy & Gaimard, 1834)
142. Species Thyone uniannulata Sluiter, 1914 accepted as Havelockia uniannulata (Sluiter, 1914)
143. Species Thyone unisemita (Stimpson, 1851) accepted as Stereoderma unisemita (Stimpson, 1851)
144. Species Thyone van Kampeni Sluiter, 1914 accepted as Havelockia vankampeni (Sluiter, 1914)
145. Species Thyone vercoi Joshua & Creed, 1915 accepted as Phyllostauros vercoi (Joshua & Creed, 1915)
146. Species Thyone wahrbergi Madsen, 1941 accepted as Thyone gadeana Perrier R., 1898 (synonym)
147. Species Thyone ceciliae Domantay, 1961 (uncertain > nomen dubium)
148. Species Thyone clarki Domantay, 1958 (uncertain > nomen dubium)
149. Species Thyone colnettensis Domantay, 1961 (uncertain > nomen dubium)
150. Species Thyone floridana Domantay, 1958 (uncertain > nomen dubium)
Subfamily Thyonellinae Panning, 1949
Genus Thyonina Thandar, 1990
1. Species Thyonina articulata (Vaney, 1908)
2. Species Thyonina bijui Thandar, 2017
3. Species Thyonina rasidae Thandar, 2017
Subfamily Thyoninae Panning, 1949
1. Genus Pseudostolus Thandar, 2021
  1. Species Pseudostolus kilberti (Rajpal & Thandar, 1999)
  2. Species Pseudostolus rapax (Koehler & Vaney, 1908)
Genus Triasemperia O'Loughlin in O'Loughlin et al., 2014
1. Species Triasemperia stola O'Loughlin in O'Loughlin et al., 2014
Genus Euphyllophorus Deichmann, 1938 accepted as Phyllophorus Grube, 1840
Genus Hemicrepis Müller, 1854 accepted as Phyllophorus Grube, 1840
Genus Holigoclados Brandt, 1835 accepted as Pentamera Ayres, 1852 (nomen oblitum)
Genus Paraphyllophorus Deichmann, 1954 accepted as Neothyonidium Deichmann, 1938
Genus Parathyone Cherbonnier, 1988 accepted as Ekmanothyone Massin, 1993
Genus Pentactes Burmeister, 1837 accepted as Cucumaria de Blainville, 1830
1. Species Pentactes andrewsii Farran, 1860 accepted as Thyone andrewsii (Farran, 1860)
Genus Semperiella Heding & Panning, 1954 accepted as Thyonidiella Heding & Panning, 1954
1. Species Semperiella drozdovi Levin & Stepanov, 1999 accepted as Thyonidiella drozdovi (Levin & Stepanov, 1999)
2. Species Semperiella tenera (Ludwig, 1875) accepted as Thyonidiella tenera (Ludwig, 1875)
Genus Subuculus Oken, 1815 accepted as Thyone Oken, 1815
Genus Thyoneria Deichmann, 1954 accepted as Stolus Selenka, 1867
Genus Thyonidiella Heding & Panning, 1954 accepted as Phyrella Heding & Panning, 1954
1. Species Thyonidiella cherbonnieri Rowe & Richmond, 2004 accepted as Phyrella ambigua (Cherbonnier, 1988)
2. Species Thyonidiella drozdovi (Levin & Stepanov, 1999) accepted as Phyrella drozdovi (Levin & Stepanov, 1999)
3. Species Thyonidiella exigua Cherbonnier, 1988 accepted as Phyllophorus exigua (Cherbonnier, 1988)
4. Species Thyonidiella kungi O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012 accepted as Phyllophorus kungi (O'Loughlin in O'Loughlin, Barmos & VandenSpiegel, 2012)
5. Species Thyonidiella oceana Heding & Panning, 1954 accepted as Phyrella fragilis (Mitsukuri & Ohshima in Ohshima, 1912)
6. Species Thyonidiella tenera (Ludwig, 1875) accepted as Phyrella tenera (Ludwig, 1875)
Genus Trachythyonidium Deichmann, 1954 accepted as Euthyonidiella Heding & Panning, 1954 (tentative subjective synonym, but seniority of name needs to be verified by publication data, as both genera were described in 1954)
Genus Urodemella Deichmann, 1944 (not available: no type species designated and post-1930 (ICZN 13.3))
1. Species Urodemella ehrenbergii (Selenka, 1868) accepted as Ohshimella ehrenbergii (Selenka, 1868)
Subfamily Urodeminae Panning, 1949 accepted as Cladolabinae Heding & Panning, 1954
Genus Isophyllophorus Liao & Pawson, 2001 represented as Phyllophorus (Isophyllophorus) Liao & Pawson, 2001
Genus Phyllonovus Cherbonnier, 1988 represented as Phyllophorus (Phyllonovus) Cherbonnier, 1988
Genus Phyllothuria Heding & Panning, 1954 represented as Phyllophorus (Phyllothuria) Heding & Panning, 1954

Environment

marine, ~~fresh~~, ~~terrestrial~~

Original description

Östergren, H. 1907. Zur Phylogenie und Systematik der Seewalzen. Särtryck ur Zoologiska Studier Tillägnade Professor T. Tullberg, pp. 191-215, Almquist & Wiksells Buchdruckerei-Aktiengesellschaft, Upsala. [details]

Descriptive notes

Taxonomic citation

WoRMS (2024). Phyllophoridae Östergren, 1907. Accessed at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=123188 on 2024-05-08

Taxonomic edit history

Date

action

2004-12-21 15:54:05Z

created

Hansson, Hans

2013-06-07 05:11:37Z

changed

Paulay, Gustav

Licensing

The webpage text is licensed under a Creative Commons Attribution 4.0 License

[taxonomic tree] [list species]

original description Östergren, H. 1907. Zur Phylogenie und Systematik der Seewalzen. Särtryck ur Zoologiska Studier Tillägnade Professor T. Tullberg, pp. 191-215, Almquist & Wiksells Buchdruckerei-Aktiengesellschaft, Upsala. [details]

context source (PeRMS) Solís-Marín, F. A.; Alvarado, J. J.; Abreu-Pérez, M.; Aguilera, O.; Alió, J.; Bacallado-Aránega, J. J.; Barraza, E.; Benavides-Serrato, M.; Benítez-Villalobos, F.; Betancourt-Fernández, L.; Borges, M.; Brandt, M.; Brogger, M. I.; Borrero-Pérez, G. H.; Buitrón-Sánchez, E.; Campos, L. S.; Cantera, J.; Clemente, S.; Cohen-Renjifo, M.; Coppard, S.; Costa-Lotufo, L. V.; del Valle-García, R.; Díaz, Y.; Díaz de Vivar, M. E.; Díaz-Martínez, J. P.; Durán-González, A.; Epherra, L.; Escolar, M.; Francisco, V.; Freire, C. A.; García-Arrarás, E.; Gil, D. G.; Guarderas, P.; Hadel, V. F.; Hearn, A.; Hernández, J. C.; Hernández-Delgado, E. A.; Herrera-Moreno, A.; Herrero-Pérezrul, M. D.; Hooker, Y.; Honey-Escandón, M. B. I.; Lodeiros, C.; Luzuriaga, M.; Manso, C. L. C.; Martín, A.; Martinez, M. I.; Martínez, S.; Moro-Abad; Mutschke, E.; Navarro, J. C.; Neira, R.; Noriega, N.; Palleiro-Nayar, J. S.; Pérez, A. F.; Pérez-Ruzafa, A.; Prieto-Rios, E.; Reyes, J.; Rodríguez, R.; Rubilar, T.; Sancho-Mejía, T.; Sangil, C.; Silva, J. R. M. C.; Sonnenholzner, J. I.; Ventura, C. R.; Tablado, A.; Tavares, Y.; Tiago, C. G.; Tuya, F.;Williams, S. M. (2013). Appendix. In: J. J. Alvarado & F. A. Solís-Marín (eds), Echinoderm Research and Diversity in Latin America. pp. 471-510. Springer; Berlin & Heidelberg. page(s): 543-654. [details]

basis of record Hansson, H.G. (2001). Echinodermata, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,. 50: pp. 336-351. (look up in IMIS) [details]

identification resource Deichmann, E. (1938). Eastern Pacific Expeditions of the New York Zoological Society. XVI. Holothurians from the western coasts of Lower California and Central America, and from the Galapagos Islands. Zoologica, New York 23: pp. 361-387. [details]

identification resource Deichmann, E. (1941). The Holothurioidea collected by the Velero III during the years1932to 1938. Part I, Dendrochirota, Allan Hancock Pacific Exped. 1932-40 Los Angeles 8: pp. 61-153. [details]

identification resource Madsen, F.J.; Hansen, B. (1994). Echinodermata Holothurioidea. Marine invertebrates of Scandinavia, 9. Scandinavian University Press: Oslo, Norway. ISBN 82-00-03936-6. 143 pp. (look up in IMIS) [details]

Present

Inaccurate

Introduced: alien

Containing type locality

Definitions

[show lower level...]

Language	Name
Japanese	グミモドキ科	[details]
Swedish	svanssjögurkor	[details]

To Biological Information System for Marine Life (BISMaL)
To European Nucleotide Archive (ENA)
To Genbank
To ITIS