Foraminifera taxon details
Profusulinella Rauzer-Chernousova & Belyaev, 1936 †
721792 (urn:lsid:marinespecies.org:taxname:721792)
accepted
Genus
Profusulinella pararhomboides Rauzer-Chernousova & Belyaev, 1936 † (type by original designation)
Kalmykoviella Ektova, 1989 † · unaccepted > junior subjective synonym (Subjective junior synonym Opinion...)
Subjective junior synonym Opinion of Ueno (2022)
- Species Profusulinella abnormis Xie, 1982 †
- Species Profusulinella acutissima Putrya, 1956 †
- Species Profusulinella albasensis van Ginkel, 1965 †
- Species Profusulinella ampla Rui, 1996 †
- Species Profusulinella apodacensis Thompson, 1948 †
- Species Profusulinella batangensis Zhang, 1982 †
- Species Profusulinella becharensis van Ginkel, 1985 †
- Species Profusulinella bella Han & Zhao, 1984 †
- Species Profusulinella beppensis Toriyama, 1958 †
- Species Profusulinella biconiformis Kireeva, 1951 †
- Species Profusulinella brevissima Putrya, 1956 †
- Species Profusulinella buensis Dalmatskaya, 1972 †
- Species Profusulinella burrensis Thompson & Shaver, 1964 †
- Species Profusulinella chaoxianensis L. Wang, 1981 †
- Species Profusulinella chernovi Rauzer-Chernousova, 1951 †
- Species Profusulinella daaoziensis Zhang, Zhou & Sheng, 2010 †
- Species Profusulinella daiyamensis Hasegawa, 1967 †
- Species Profusulinella decora Thompson, 1948 †
- Species Profusulinella deplanata Dzhenchuraeva, 1979 †
- Species Profusulinella deprati (Beede & Kniker, 1924) †
- Species Profusulinella diminutiva (Thompson, 1936) †
- Species Profusulinella dissidens Han & Zhao, 1984 †
- Species Profusulinella dubia Villa Otero, 1995 †
- Species Profusulinella duijiugouensis Zhuang, 1990 †
- Species Profusulinella elegans Saltovskaya, 1974 †
- Species Profusulinella ellipsoidalis Rumyantseva, 1974 †
- Species Profusulinella elliptica Chen J.R., 1978 †
- Species Profusulinella eolibrovichi Safonova, 1951 †
- Species Profusulinella eoprisca Ivanova, 2000 †
- Species Profusulinella extensa Rauzer-Chernousova, 1961 †
- Species Profusulinella faqianensis Zhang, Zhou & Sheng, 2010 †
- Species Profusulinella fenghuangshanensis L. Wang, 1981 †
- Species Profusulinella fittsi (Thompson, 1935) †
- Species Profusulinella fluxoidalis Manukalova, 1950 †
- Species Profusulinella fukujiensis Igo, 1957 †
- Species Profusulinella fusiformis Sada, 1972 †
- Species Profusulinella gaokouwuensis Qian, 1982 †
- Species Profusulinella grozdilovae Liêm, 1967 †
- Species Profusulinella grozdilovae Ektova, 1972 † (accepted > unreplaced junior homonym, Primary junior homonym of Profusulinella grozdilovae Liêm, 1967)
- Species Profusulinella guangnanensis Chen, Yang, Wang & Zhang, 1991 †
- Species Profusulinella hanshanensis Qian, 1982 †
- Species Profusulinella hayasakai (Watanabe, 1974) †
- Species Profusulinella integra Chernova, 1961 †
- Species Profusulinella intermedia Bensh, 1969 †
- Species Profusulinella jamnaensis Ivanova, 2000 †
- Species Profusulinella jifukensis Da in Da & Sun, 1983 †
- Species Profusulinella jinghensis Da in Da & Sun, 1983 †
- Species Profusulinella jomdaensis Chen J.R., 1978 †
- Species Profusulinella keramiliensis Sun in Da & Sun, 1983 †
- Species Profusulinella kireevae Dzhenchuraeva, 1979 †
- Species Profusulinella koxkudukensis Da in Da & Sun, 1983 †
- Species Profusulinella latispiralis Safonova, 1951 †
- Species Profusulinella liaoningensis Han, 1980 †
- Species Profusulinella longitica Manukalova, 1956 †
- Species Profusulinella majiaobaensis Chen J.R., 1978 †
- Species Profusulinella manuensis Roberts, 1949 †
- Species Profusulinella maopanshanensis Liu, Xiao & Dong, 1978 †
- Species Profusulinella marblensis Thompson, 1947 †
- Species Profusulinella martinezi Villa, 2016 †
- Species Profusulinella melekessensis Dalmatskaya, 1972 †
- Species Profusulinella millcreekensis Douglass & Nestell, 1984 †
- Species Profusulinella minima Dzhenchuraeva, 1979 †
- Species Profusulinella munda Thompson, 1948 †
- Species Profusulinella oblonga Potievskaya, 1964 †
- Species Profusulinella ohioensis Douglass, 1987 †
- Species Profusulinella orbicularis Bensh, 1969 †
- Species Profusulinella ovataeformis Dalmatskaya, 1972 †
- Species Profusulinella parafittsi Rauzer-Chernousova & Safonova, 1951 †
- Species Profusulinella paranibelensis Chang, 1960 †
- Species Profusulinella paraprisca Bensh, 1969 †
- Species Profusulinella pararhomboides Rauzer-Chernousova & Belyaev, 1936 †
- Species Profusulinella paratimanica Rauzer-Chernousova, 1951 †
- Species Profusulinella parva (Lee & Chen, 1930) †
- Species Profusulinella plummeri Skinner & Wilde, 1954 †
- Species Profusulinella polasnensis Safonova, 1951 †
- Species Profusulinella posadai Villa Otero, 1995 †
- Species Profusulinella postpararhomboides Dzhenchuraeva, 1979 †
- Species Profusulinella praelibrovichi Safonova, 1951 †
- Species Profusulinella praeovata Manukalova-Grebenyuk, Ilyina & Serezhnikova, 1969 †
- Species Profusulinella praeprisca Solovieva, 1977 †
- Species Profusulinella prima Lebedeva, 1966 †
- Species Profusulinella primaeva (Skinner, 1931) †
- Species Profusulinella primitiva Sosnina in Grozdilova & Lebedeva, 1954 †
- Species Profusulinella probiconica Watanabe, 1974 †
- Species Profusulinella propria Nikolaev, 2011 †
- Species Profusulinella pseudorhomboides Putrya in Putrya & Leontovich, 1948 †
- Species Profusulinella quasiparva Rui, 1983 †
- Species Profusulinella quasirhomboides Liu, Xiao & Dong, 1978 †
- Species Profusulinella quydatensis Liêm, 1967 †
- Species Profusulinella regia Thompson, 1948 †
- Species Profusulinella rhombiformis Brazhnikova & Potievskaya, 1948 †
- Species Profusulinella rhomboides (Lee & Chen, 1930) †
- Species Profusulinella rotundata Putrya, 1956 †
- Species Profusulinella sitteri van Ginkel, 1987 †
- Species Profusulinella spicata Thompson, 1948 †
- Species Profusulinella springensis Marshall, 1969 †
- Species Profusulinella subsphaerica Chen J.R., 1978 †
- Species Profusulinella talimuensis Han & Zhao, 1984 †
- Species Profusulinella tchauwaica Dzhenchuraeva, 1979 †
- Species Profusulinella terskeica Dzhenchuraeva, 1979 †
- Species Profusulinella thompsoni Clopine, 1992 †
- Species Profusulinella tianshanensis Da in Da & Sun, 1983 †
- Species Profusulinella tonkinensis Saurin, 1970 †
- Species Profusulinella topiliensis (Putrya, 1938) †
- Species Profusulinella trisulcata (Thompson, 1935) †
- Species Profusulinella turkestanica Dzhenchuraeva, 1979 †
- Species Profusulinella urumqiensis Da in Da & Sun, 1983 †
- Species Profusulinella walnutensis Ross & Sabins, 1965 †
- Species Profusulinella weiningica Zhang, 1974 †
- Species Profusulinella whitetailensis Ross & Sabins, 1965 †
- Species Profusulinella yazitangica Zhang, Zhou & Sheng, 2010 †
- Species Profusulinella yunnanica Chen, Yang, Wang & Zhang, 1991 †
- Species Profusulinella zhanatanensis Nikolaev, 2011 †
- Species Profusulinella zhaoudaensis Han, 1980 †
- Species Profusulinella zhongzanica Zhang, 1982 †
- Species Profusulinella zhuangzishangensis Zhang, 1982 †
- Species Profusulinella chiktongensis Kang, 1970 † (uncertain > unassessed)
- Subgenus Profusulinella (Depratina) Leven, 2009 † accepted as Depratina Solovieva in Rauzer-Chernousova et al., 1996 † (Nomen translatum)
- Species Profusulinella (Depratina) pseudoparva Leven, Davydov & Gorgij, 2006 †
- Species Profusulinella (Depratina) beppensis Toriyama, 1958 † accepted as Profusulinella beppensis Toriyama, 1958 † (Opinion of Kobayashi (2017))
- Species Profusulinella (Depratina) omiensis Watanabe, 1974 † accepted as Solovievaia omiensis (Watanabe, 1974) †
- Species Profusulinella (Depratina) parva (Lee & Chen, 1930) † accepted as Profusulinella parva (Lee & Chen, 1930) †
- Species Profusulinella (Depratina) subovata Safonova in Rauzer-Chernousova et al., 1951 † accepted as Solovievaia subovata (Safonova in Rauzer-Chernousova et al., 1951) †
- Species Profusulinella (Depratina) timanica Kireeva, 1951 † accepted as Depratina timanica (Kireeva, 1951) †
- Subgenus Profusulinella (Taitzehoella) A.D. Miklukho-Maklay et al., 1959 † accepted as Taitzehoella Sheng, 1951 † (Opinion of Loeblich & Tappan, 1987 Nomen translatum)
- Species Profusulinella aljutovica Rauzer-Chernousova, 1938 † accepted as Aljutovella aljutovica (Rauzer-Chernousova, 1938) † (Type species of Aljutovella)
- Species Profusulinella arta Leontovich in Rauzer-Chernousova et al., 1951 † accepted as Solovievaia arta (Leontovich in Rauzer-Chernousova et al., 1951) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella bedakensis Solovieva, 1977 † accepted as Moellerites bedakensis (Solovieva, 1977) † (Opinion of Solovieva (1986))
- Species Profusulinella bisyllaba Ektova, 1972 † accepted as Staffellaeformes bisyllaba (Ektova, 1972) † (Opinion of Solovieva (1986))
- Species Profusulinella bona Grozdilova & Lebedeva, 1954 † accepted as Staffellaeformes bona (Grozdilova & Lebedeva, 1954) † (Opinion of Solovieva (1986))
- Species Profusulinella carasaica Dzhenchuraeva, 1979 † accepted as Solovievaia carasaica (Dzhenchuraeva, 1979) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella cavis Dalmatskaya, 1961 † accepted as Solovievaia cavis (Dalmatskaya, 1961) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella chaohuensis L. Wang, 1981 † accepted as Solovievaia chaohuensis (L. Wang, 1981) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella compacta Rumyantseva, 1974 † accepted as Solovievaia compacta (Rumyantseva, 1974) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella constans Safonova in Rauzer-Chernousova et al., 1951 † accepted as Solovievaia constans (Safonova in Rauzer-Chernousova et al., 1951) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella convoluta (Lee & Chen, 1930) † accepted as Depratina convoluta (Lee & Chen, 1930) † (Opinion of Akbas and Okuyucu (2021))
- Species Profusulinella copiosa Thompson, 1948 † accepted as Staffellaeformes copiosa (Thompson, 1948) † accepted as Solovievaia copiosa (Thompson, 1948) † (unaccepted > superseded combination, Opinion of Solovieva (1986))
- Species Profusulinella dagmarae (Safonova, 1951) † accepted as Aljutovella dagmarae Safonova, 1951 † (unaccepted > superseded combination)
- Species Profusulinella exempla (Stewart, 1958) † accepted as Frumentella exempla Stewart, 1958 † (unaccepted > superseded combination, Opinion of Wilde (2006))
- Species Profusulinella hinodensis Kobayashi, 1994 † accepted as Solovievaia hinodensis (Kobayashi, 1994) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella kentuckyensis Thompson & Riggs, 1959 † accepted as Moellerites kentuckyensis (Thompson & Riggs, 1959) † (Opinion of Solovieva (1986))
- Species Profusulinella kungesaica Bogush & Yuferev, 1976 † accepted as Staffellaeformes kungesaica (Bogush & Yuferev, 1976) † (unaccepted > superseded combination, Opinion of Kobayashi and Vachard (2022))
- Species Profusulinella librovitchi (Dutkevich, 1934) † accepted as Taitzehoella librovitchi (Dutkevich, 1934) † (Opinion of Leven (2009))
- Species Profusulinella montichomata Sheng & Wang, 1976 † accepted as Solovievaia montichomata (Sheng & Wang, 1976) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella muruntavica Rumyantseva, 1974 † accepted as Solovievaia muruntavica (Rumyantseva, 1974) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella mutabilis Safonova, 1951 † accepted as Taitzehoella mutabilis (Safonova, 1951) † (Opinion of Leven (2009))
- Species Profusulinella nuratavensis Solovieva, 1977 † accepted as Solovievaia nuratavensis (Solovieva, 1977) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella obscura (Lee & Chen, 1930) † accepted as Schubertina obscura (Lee & Chen, 1930) †
- Species Profusulinella omiensis Watanabe, 1974 † accepted as Solovievaia omiensis (Watanabe, 1974) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella ovata Rauzer-Chernousova, 1938 † accepted as Ovatella ovata (Rauzer-Chernousova, 1938) † accepted as Solovievaia ovata (Rauzer-Chernousova, 1938) † (Type species of Ovatella)
- Species Profusulinella paraaljutovica (Safonova, 1951) † accepted as Aljutovella paraaljutovica Safonova, 1951 † (unaccepted > superseded combination)
- Species Profusulinella parasaratovica (Safonova, 1951) † accepted as Aljutovella parasaratovica Safonova, 1951 † (unaccepted > superseded combination, Opinion of Isakova (2012))
- Species Profusulinella paraschubertellinoides Putrya in Putrya & Leontovich, 1948 † accepted as Fusulinella paraschubertellinoides (Putrya in Putrya & Leontovich, 1948) † (Opinion of Kahler and Kahler (1966), Rozovskaya (1975))
- Species Profusulinella parasimplex Bensh, 1969 † accepted as Moellerites parasimplex (Bensh, 1969) † (unaccepted > superseded combination, Opinion of Orlov-Labkovsky and Bensh (2015))
- Species Profusulinella postaljutovica (Safonova, 1951) † accepted as Priscoidella postaljutovica (Safonova, 1951) †
- Species Profusulinella praecursor (Deprat, 1913) † accepted as Solovievaia praecursor (Deprat, 1913) †
- Species Profusulinella primitiva Sosnina in Ektova, 1976 † accepted as Profusulinella primitiva Sosnina in Grozdilova & Lebedeva, 1954 † (Objective synonym)
- Species Profusulinella prisca (Deprat, 1912) † accepted as Depratina prisca (Deprat, 1912) † (Type species of Depratina)
- Species Profusulinella priscoidea Rauzer-Chernousova, 1938 † accepted as Priscoidella priscoidea (Rauzer-Chernousova, 1938) † (Type species of Priscoidella)
- Species Profusulinella prolibrovichi Rauzer-Chernousova, 1951 † accepted as Taitzehoella prolibrovichi (Rauzer-Chernousova, 1951) † (unaccepted > superseded combination)
- Species Profusulinella pseudoaljutovica (Rauzer-Chernousova, 1951) † accepted as Tikhonovichiella pseudoaljutovica (Rauzer-Chernousova, 1951) † (unaccepted > superseded combination)
- Species Profusulinella pseudolibrovichi Safonova, 1951 † accepted as Taitzehoella pseudolibrovichi (Safonova, 1951) † (Opinion of Villa et al. (2021))
- Species Profusulinella pseudoobscura (Chen, 1934) † accepted as Schubertella pseudoobscura Chen, 1934 † (Opinion of Igo (1996))
- Species Profusulinella pseudoparva Leven, Davydov & Gorgij, 2006 † accepted as Profusulinella (Depratina) pseudoparva Leven, Davydov & Gorgij, 2006 † (Opinion of Leven (2009))
- Species Profusulinella saratovica Putrya in Putrya & Leontovich, 1948 † accepted as Aljutovella saratovica (Putrya in Putrya & Leontovich, 1948) † (Opinion of Kahler and Kahler (1966), Rozovskaya (1975))
- Species Profusulinella simplex Safonova, 1951 † accepted as Solovievaia simplex (Safonova, 1951) † (unaccepted > superseded combination, Opinion of Akbas and Okuyucu (2022), Vachard (2023) pers. comm. [transitional form to Moellerites and later to Fusulinella])
- Species Profusulinella skelnevatica Putrya in Putrya & Leontovich, 1948 † accepted as Skelnevatella skelnevatica (Putrya in Putrya & Leontovich, 1948) † (Type species of Skelnevatella)
- Species Profusulinella sokolensis Ivanova, 2000 † accepted as Solovievaia sokolensis (Ivanova, 2000) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella staffellaeformis Kireeva, 1951 † accepted as Staffellaeformes staffellaeformis (Kireeva, 1951) † (Type species of Staffellaeformes )
- Species Profusulinella staffelloides Manukalova, 1948 † accepted as Staffellaeformes staffelloides (Manukalova, 1948) † (Opinion of Solovieva (1986))
- Species Profusulinella subovata Safonova in Rauzer-Chernousova et al., 1951 † accepted as Solovievaia subovata (Safonova in Rauzer-Chernousova et al., 1951) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella syzranica Rauzer-Chernousova, 1951 † accepted as Taitzehoella syzranica (Rauzer-Chernousova, 1951) † (Opinion of Zhang et al. (2010))
- Species Profusulinella tashliensis Lebedeva in Stepanov et al., 1975 † accepted as Staffellaeformes tashliensis (Lebedeva in Stepanov et al., 1975) † (Opinion of Solovieva (1986))
- Species Profusulinella tchotchiai Grozdilova & Lebedeva, 1960 † accepted as Solovievaia tchotchiai (Grozdilova & Lebedeva, 1960) † (Opinion of Vachard and Le Coze (2018))
- Species Profusulinella toriyamai Sada, 1961 † accepted as Staffellaeformes toriyamai (Sada, 1961) † accepted as Solovievaia toriyamai (Sada, 1961) † (unaccepted > superseded combination, Opinion of Solovieva (1986))
- Species Profusulinella wangyui Sheng, 1958 † accepted as Dagmarella wangyui (Sheng, 1958) †
marine, brackish, fresh, terrestrial
fossil only
feminine
Rauzer-Chernousova, D. M.; Belyaev, G.; Reitlinger, E. (1936). Верхнепалеозойские фораминиферы Печорского края - Upper Paleozoic foraminifera of the Pechora region. <em>Труды полярной комиссии АН СССР - Proceedings of the Polar Commission AS USSR.</em> 28: 159–232.
page(s): p. 175; note: p. 220 (German)) [details] Available for editors
[request]
page(s): p. 175; note: p. 220 (German)) [details] Available for editors
Description Description: Test minute to medium sized, inflated fusiform to subellipsoidal, with bluntly rounded poles. First whorl...
Diagnosis Test small, up to about 1 mm in length, fusiform, globular proloculus followed by closely coiled early whorls, rapidly...
Description Description: Test minute to medium sized, inflated fusiform to subellipsoidal, with bluntly rounded poles. First whorl subdiscoidal to spherical and occasionally coiled at large angles to the coiling axis. Coiling uniform but rather loose. Septa are thin. Septal fluting absent or poorly developed in polar regions. Proloculus spherical. Tunnel rather narrow. Chomata massive and wide. Wall unilayered, dark microgranular. Aperture terminal, simple.
Occurrence: Late Bashkirian-early Moscovian; probably cosmopolitan. Biozones 10 (Tashastian) to 12 (Vereian) of Akiyoshi area (this study).
(Kobayashi and Vachard (2022)). [details]
Occurrence: Late Bashkirian-early Moscovian; probably cosmopolitan. Biozones 10 (Tashastian) to 12 (Vereian) of Akiyoshi area (this study).
(Kobayashi and Vachard (2022)). [details]
Diagnosis Test small, up to about 1 mm in length, fusiform, globular proloculus followed by closely coiled early whorls, rapidly...
Diagnosis Test small, up to about 1 mm in length, fusiform, globular proloculus followed by closely coiled early whorls, rapidly becoming elongate after a sharp change in coiling axis, septa plane and not fluted; wall with three layers, tectum, diaphanotheca, and upper tectorium, with lower tectorium in a few chambers of some advanced species, chomata prominent, asymmetrical, secondary thickening extending laterally toward the poles. U. Carboniferous (Moscovian); USA: Texas; Peru; China; Japan; USSR. (Loeblich & Tappan, 1987, Foraminiferal Genera and Their Classification) [details]
Hayward, B.W.; Le Coze, F.; Vachard, D.; Gross, O. (2021). World Foraminifera Database. Profusulinella Rauzer-Chernousova & Belyaev, 1936 †. Accessed at: http://www.marinespecies.org/foraminifera/aphia.php?p=taxdetails&id=721792 on 2026-07-18
Date
action
by
original description
Rauzer-Chernousova, D. M.; Belyaev, G.; Reitlinger, E. (1936). Верхнепалеозойские фораминиферы Печорского края - Upper Paleozoic foraminifera of the Pechora region. <em>Труды полярной комиссии АН СССР - Proceedings of the Polar Commission AS USSR.</em> 28: 159–232.
page(s): p. 175; note: p. 220 (German)) [details] Available for editors
[request]
original description (of Kalmykoviella Ektova, 1989 †) Ektova, L. A. (1989). Новое семейство Fusiellidae и генетические связи некоторых среднекаменноугольных фузулинид - A new family of Fusiellidae and genetic bonds of some Middle Carboniferous fusulinids. <em>Annual of the all-union Paleontological Society.</em> 22: 43-50.
page(s): p. 47 footnote; note: Originally included in the Schubertellidae. [details] Available for editors
[request]
basis of record Loeblich, A. R.; Tappan, H. (1987). Foraminiferal Genera and their Classification. Van Nostrand Reinhold Company, New York. 970pp., available online at https://books.google.pt/books?id=n_BqCQAAQBAJ [details] Available for editors
[request]
page(s): p. 175; note: p. 220 (German)) [details] Available for editors
original description (of Kalmykoviella Ektova, 1989 †) Ektova, L. A. (1989). Новое семейство Fusiellidae и генетические связи некоторых среднекаменноугольных фузулинид - A new family of Fusiellidae and genetic bonds of some Middle Carboniferous fusulinids. <em>Annual of the all-union Paleontological Society.</em> 22: 43-50.
page(s): p. 47 footnote; note: Originally included in the Schubertellidae. [details] Available for editors
basis of record Loeblich, A. R.; Tappan, H. (1987). Foraminiferal Genera and their Classification. Van Nostrand Reinhold Company, New York. 970pp., available online at https://books.google.pt/books?id=n_BqCQAAQBAJ [details] Available for editors
From editor or global species database
Description Description: Test minute to medium sized, inflated fusiform to subellipsoidal, with bluntly rounded poles. First whorl subdiscoidal to spherical and occasionally coiled at large angles to the coiling axis. Coiling uniform but rather loose. Septa are thin. Septal fluting absent or poorly developed in polar regions. Proloculus spherical. Tunnel rather narrow. Chomata massive and wide. Wall unilayered, dark microgranular. Aperture terminal, simple.Occurrence: Late Bashkirian-early Moscovian; probably cosmopolitan. Biozones 10 (Tashastian) to 12 (Vereian) of Akiyoshi area (this study).
(Kobayashi and Vachard (2022)). [details]
Diagnosis Test small, up to about 1 mm in length, fusiform, globular proloculus followed by closely coiled early whorls, rapidly becoming elongate after a sharp change in coiling axis, septa plane and not fluted; wall with three layers, tectum, diaphanotheca, and upper tectorium, with lower tectorium in a few chambers of some advanced species, chomata prominent, asymmetrical, secondary thickening extending laterally toward the poles. U. Carboniferous (Moscovian); USA: Texas; Peru; China; Japan; USSR. (Loeblich & Tappan, 1987, Foraminiferal Genera and Their Classification) [details]