original descriptionBenham, W.B. (1886). Studies on Earthworms II. <em>Quarterly Journal of Microscopical Science.</em> 27 (105):77-108 + Plate VIII-IX., available online athttps://www.biodiversitylibrary.org/page/13819623 page(s): 94 [details]
additional sourceBeddard, F.E. (1895). A monograph of the order of Oligochaeta. <em>[Book].</em> 769 pp. The Clarendon Press, Oxford., available online athttps://www.biodiversitylibrary.org/item/71343 page(s): 558 [details]