Traits taxon details
original description
Beddard, F. E. (1892). XVII.—The earthworms of the Vienna Museum. <em>Journal of Natural History.</em> 9(50): 113-134. page(s): 128 + Plate VII, fig 4-5 [details]
additional source
Beddard, F.E. (1895). A monograph of the order of Oligochaeta. <em>[Book].</em> 769 pp. The Clarendon Press, Oxford., available online at https://www.biodiversitylibrary.org/item/71343 page(s): 663 [details]
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