Thiercelin, N.; Schubart, C.D. (2014). Transisthmian differentiation in the tree-climbing mangrove crab Aratus H. Milne Edwards, 1853 (Crustacea, Brachyura, Sesarmidae), with description of a new species from the tropical eastern Pacific. Zootaxa. 3793(5): 545-560.
Transisthmian differentiation in the tree-climbing mangrove crab <em>Aratus </em>H. Milne Edwards, 1853 (Crustacea, Brachyura, Sesarmidae), with description of a new species from the tropical eastern Pacific