Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 221, Fig. 10: U. Devonian (Frasnian), Saratov Oblast, Russian Platform. Holotype, x 120 (from E. V. Bykova, 1952).