Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 342, Figs. 28-33: M. Miocene (U. Tortonian). Ukraine. 28-30, opposite sides and apertural view of holotype, x 40; 31-33, ontogenetic development of apertural tooth (from Serova, 1961).