Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 39, Figs. 17-20: Paleocene, Bukhara, Uzbekistan, side views, X 100; 18, holotype; 20, U. Eocene, Amu Darya Basin, Uzbekistan, side view, x 100 (from Suleymanov, 1960).