Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 220, Figs. 5-6: Devonian, Russia. 5, U. Devonian (Frasnian), Volgograd (Stalingrad) Oblast, holotype, x 100; 6, M. Devonian (Givetian), Voronezh Oblast, specimen with early chambers preserved, x 100 (from E. V. Bykova, 1955).