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Penna, A., Fraga, S., Maso, M., Giacobbe, M. G., Bravo, I., Garces, E., Vila, M., Bertozzini, E., Andreoni, F., Luglie, A. & Vernesi, C. 2008. Phylogenetic relationships among the Mediterranean Alexandrium (Dinophyceae) species based on sequences of 5.8S gene and Internal Transcript Spacers of the rRNA operon. European Journal of Phycology 43:163-78.