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Manuel, M.; Borchiellini, C.; Alivon, E.; Le Parco, Y.; Vacelet, J.; Boury-Esnault, N. (2003). Phylogeny and Evolution of Calcareous Sponges: Monophyly of Calcinea and Calcaronea, High Level of Morphological Homoplasy, and the Primitive Nature of Axial Symmetry. Systematic Biology. 53, 311-333.