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Kritsky, D. C.; Boeger, W. A.; Jégu, M. (1997). Neotropical Monogenoidea. 29. Ancyrocephalinae (Dactylogyridae) of piranha and their relatives (Teleostei, Serrasalmidae) from Brazil: Species of Amphithecium Boeger and Kritsky, 1988, Heterothecium gen. n. and Pithanothecium gen. n. Journal of the Helminthological Society of Washington. 64, 25-54.