WoRMS taxon details
Varunidae H. Milne Edwards, 1853
- Subfamily Asthenognathinae Stimpson, 1858
- Genus Asthenognathus Stimpson, 1858
- Species Asthenognathus alleronensis Pasini, Garassino & De Angeli, 2017 †
- Species Asthenognathus atlanticus Monod, 1933
- Species Asthenognathus australensis Feldmann, Schweitzer, Casadío & Griffin, 2011 †
- Species Asthenognathus cornishorum Schweitzer & Feldmann, 1999 †
- Species Asthenognathus fernandezi Ferratges, Zamora & Aurell, 2022 †
- Species Asthenognathus globosa (Karasawa, 1990) †
- Species Asthenognathus hexagonum Rathbun, 1909
- Species Asthenognathus inaequipes Stimpson, 1858
- Species Asthenognathus laverdensis De Angeli & Garassino, 2006 †
- Species Asthenognathus microspinus Casadío, De Angeli, Feldmann, Garassino, Hetler, Parras & Schweitzer, 2004 †
- Species Asthenognathus rakosensis Müller, 2006 †
- Species Asthenognathus sakumotoi Karasawa, 2018 †
- Species Asthenognathus urretae Schweitzer & Feldmann, 2001 †
- Species Asthenognathus gallardoi Serène & Soh, 1976 accepted as Gopkittisak gallardoi (Serène & Soh, 1976) (unaccepted > superseded combination)
- Genus Globihexapus Schweitzer & Feldmann, 2001 †
- Tribe Gustavini Števčić, 2011 accepted as Aphanodactylidae Ahyong & Ng, 2009
- Genus Asthenognathus Stimpson, 1858
- Subfamily Cyclograpsinae H. Milne Edwards, 1853
- Genus Austrohelice Sakai, Türkay & Yang, 2006
- Genus Chasmagnathus De Haan, 1833
- Species Chasmagnathus convexus (De Haan, 1835)
- Species Chasmagnathus convexus Haswell, 1881 accepted as Helograpsus haswellianus (Whitelegge, 1890) (unaccepted > junior homonym)
- Species Chasmagnathus georgei Clark, 1987 † accepted as Parahelice georgei (Clark, 1987) (unaccepted > superseded combination)
- Species Chasmagnathus granulatus Dana, 1851 accepted as Neohelice granulata (Dana, 1851)
- Species Chasmagnathus Haswellianus Whitelegge, 1890 accepted as Helograpsus haswellianus (Whitelegge, 1890) (unaccepted > superseded combination)
- Species Chasmagnathus laevis Dana, 1851 accepted as Paragrapsus laevis (Dana, 1851) (unaccepted > superseded combination)
- Species Chasmagnathus subquadratus Dana, 1851 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > superseded combination)
- Genus Cyclograpsus H. Milne Edwards, 1837
- Species Cyclograpsus audouinii H. Milne Edwards, 1837
- Species Cyclograpsus barbatus (MacLeay, 1838)
- Species Cyclograpsus beccarii Nobili, 1899
- Species Cyclograpsus cinereus Dana, 1851
- Species Cyclograpsus directus Karasawa, 1989 †
- Species Cyclograpsus escondidensis Rathbun in Glassell, 1933
- Species Cyclograpsus eydouxi H. Milne Edwards, 1853
- Species Cyclograpsus granulatus Dana, 1851
- Species Cyclograpsus granulosus H. Milne Edwards, 1853
- Species Cyclograpsus henshawi Rathbun, 1902
- Species Cyclograpsus incisus Shen, 1940
- Species Cyclograpsus insularum Campbell & Griffin, 1966
- Species Cyclograpsus integer H. Milne Edwards, 1837
- Species Cyclograpsus intermedius Ortmann, 1894
- Species Cyclograpsus lavauxi H. Milne Edwards, 1853
- Species Cyclograpsus longipes Stimpson, 1858
- Species Cyclograpsus lucidus Dai in Dai, Yang, Song & Chen, 1986
- Species Cyclograpsus lwangi Hu & Tao, 1999 †
- Species Cyclograpsus minutus Hombron & Jacquinot, 1846
- Species Cyclograpsus pumilio Hangai & Fukui in Hangai, Kitaura, Wada & Fukui, 2009
- Species Cyclograpsus punctatus H. Milne Edwards, 1837
- Species Cyclograpsus rectangularis Karasawa, 1989 †
- Species Cyclograpsus sanctaecrucis Griffin, 1968
- Species Cyclograpsus unidens Nobili, 1905
- Species Cyclograpsus audini H. Milne Edwards, 1837 accepted as Cyclograpsus audouinii H. Milne Edwards, 1837 (unaccepted > unjustified emendation)
- Species Cyclograpsus crenulatus H. Milne Edwards, 1837 accepted as Hemigrapsus crenulatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus gaimardii H. Milne Edwards, 1837 accepted as Paragrapsus gaimardii (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus gibbus Hombron & Jacquinot, 1846 accepted as Hemigrapsus gibbus (Hombron & Jacquinot, 1846) (unaccepted > superseded combination)
- Species Cyclograpsus gnatherion Kinahan, 1857 accepted as Pinnotherelia laevigata H. Milne Edwards & Lucas, 1843 (unaccepted > junior subjective synonym)
- Species Cyclograpsus laevis Hess, 1865 accepted as Cyclograpsus audouinii H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Species Cyclograpsus latreillii H. Milne Edwards, 1837 accepted as Pseudohelice latreillii (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus lophopus Nobili, 1906 accepted as Cyclorma lophopum (Nobili, 1906)
- Species Cyclograpsus marmoratus White, 1847 (unaccepted > nomen nudum)
- Species Cyclograpsus occidentalis A. Milne-Edwards, 1878 accepted as Cyclograpsus integer H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Species Cyclograpsus octodentatus H. Milne Edwards, 1837 accepted as Leptograpsodes octodentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus parvulus De Man, 1896 accepted as Cyclograpsus integer H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Variety Cyclograpsus parvulus var. unidens Nobili, 1905 accepted as Cyclograpsus unidens Nobili, 1905 (unaccepted > superseded combination)
- Species Cyclograpsus quadridentatus H. Milne Edwards, 1837 accepted as Paragrapsus quadridentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus reynaudi H. Milne Edwards, 1853 accepted as Cyclograpsus punctatus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Species Cyclograpsus sexdentatus H. Milne Edwards, 1837 accepted as Hemigrapsus sexdentatus (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Cyclograpsus whitei H. Milne Edwards, 1853 accepted as Cyclograpsus lavauxi H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
- Species Cyclograpsus tasmanicus Hombron & Jacquinot, 1846 (uncertain > taxon inquirendum)
- Genus Helicana Sakai & Yatsuzuka, 1980
- Genus Helice De Haan, 1833
- Species Helice epicure Ng, Naruse & Shih, 2018
- Species Helice formosensis Rathbun, 1931
- Species Helice latimera Parisi, 1918
- Species Helice tientsinensis Rathbun, 1931
- Species Helice tridens (De Haan, 1835)
- Subspecies Helice tridens formosensis Rathbun, 1931 accepted as Helice formosensis Rathbun, 1931 (unaccepted > superseded combination)
- Subspecies Helice tridens pingi Rathbun, 1931 accepted as Helice latimera Parisi, 1918 (unaccepted > junior subjective synonym)
- Subspecies Helice tridens sheni Sakai, 1939 accepted as Helicana wuana (Rathbun, 1931) (unaccepted > junior subjective synonym)
- Subspecies Helice tridens tientsinensis Rathbun, 1931 accepted as Helice tientsinensis Rathbun, 1931 (unaccepted > superseded combination)
- Variety Helice tridens var. latimera Parisi, 1918 accepted as Helice latimera Parisi, 1918 (unaccepted > superseded combination)
- Subspecies Helice tridens wuana Rathbun, 1931 accepted as Helice wuana Rathbun, 1931 (unaccepted > superseded combination)
- Subgenus Helice (Helicana) Sakai & Yatsuzuka, 1980 accepted as Helicana Sakai & Yatsuzuka, 1980 (status change)
- Species Helice (Helicana) japonica Sakai & Yatsuzuka, 1980 accepted as Helicana japonica (Sakai & Yatsuzuka, 1980) (unaccepted > superseded combination)
- Species Helice crassa Dana, 1851 accepted as Austrohelice crassa (Dana, 1851) (unaccepted > superseded combination)
- Species Helice dentipes Heller, 1865 accepted as Metaplax dentipes (Heller, 1865) (unaccepted > superseded combination)
- Species Helice gaudichaudi H. Milne Edwards, 1853 accepted as Neohelice granulata (Dana, 1851) (unaccepted > junior subjective synonym)
- Species Helice japonica Sakai & Yatsuzuka, 1980 accepted as Helicana japonica (Sakai & Yatsuzuka, 1980) (unaccepted > superseded combination)
- Species Helice latreillei (H. Milne Edwards, 1837) accepted as Helice tridens (De Haan, 1835) (unaccepted > junior subjective synonym, unjustified emendation)
- Species Helice latreilli (H. Milne Edwards, 1837) accepted as Helice tridens (De Haan, 1835) (unaccepted > junior subjective synonym, unjustified emendation)
- Species Helice leachi Hess, 1865 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > misspelling - incorrect subsequent spelling)
- Species Helice Leachii Hess, 1865 accepted as Pseudohelice subquadrata (Dana, 1851) (unaccepted > junior subjective synonym)
- Species Helice lucasi H. Milne Edwards, 1853 accepted as Austrohelice crassa (Dana, 1851) (unaccepted > junior subjective synonym)
- Species Helice pilimana A. Milne-Edwards, 1873 accepted as Parahelice pilimana (A. Milne-Edwards, 1873) (unaccepted > superseded combination)
- Species Helice spinicarpa H. Milne Edwards, 1853 accepted as Chasmagnathus convexus (De Haan, 1835) (unaccepted > junior subjective synonym)
- Species Helice wuana Rathbun, 1931 accepted as Helicana wuana (Rathbun, 1931) (unaccepted > superseded combination)
- Genus Helograpsus Campbell & Griffin, 1966
- Genus Metaplax H. Milne Edwards, 1852
- Species Metaplax crenulata (Gerstaecker, 1856)
- Species Metaplax dentipes (Heller, 1865)
- Species Metaplax distincta H. Milne Edwards, 1852
- Species Metaplax elegans De Man, 1888
- Species Metaplax gocongensis Davie & V.X. Nguyen, 2003
- Species Metaplax indica H. Milne Edwards, 1852
- Species Metaplax intermedia De Man, 1888
- Species Metaplax longipes Stimpson, 1858
- Species Metaplax occidentalis Pretzmann, 1971
- Species Metaplax sheni Gordon, 1930
- Species Metaplax tredecim Tweedie, 1950
- Species Metaplax crassipes De Man, 1892 accepted as Metaplax elegans De Man, 1888 (unaccepted > junior subjective synonym)
- Species Metaplax distinctus H. Milne Edwards, 1852 accepted as Metaplax distincta H. Milne Edwards, 1852 (unaccepted > incorrect grammatical agreement of specific epithet)
- Species Metaplax hirsutimana Grant & McCulloch, 1906 accepted as Australoplax tridentata (A. Milne-Edwards, 1873) (unaccepted > junior subjective synonym)
- Species Metaplax hirtipes Heller, 1862 accepted as Hemiplax hirtipes (Hombron & Jacquinot, 1846) (unaccepted > junior subjective synonym)
- Species Metaplax indicus H. Milne Edwards, 1852 accepted as Metaplax indica H. Milne Edwards, 1852 (unaccepted > incorrect grammatical agreement of specific epithet)
- Subspecies Metaplax indicus occidentalis Pretzmann, 1971 accepted as Metaplax occidentalis Pretzmann, 1971 (unaccepted > superseded combination)
- Species Metaplax intermedius De Man, 1888 accepted as Metaplax intermedia De Man, 1888 (unaccepted > superseded combination)
- Species Metaplax takahashii Sakai, 1939 accepted as Metaplax longipes Stimpson, 1858 (unaccepted > junior subjective synonym)
- Genus Miosesarma Karasawa, 1989 †
- Genus Neohelice Sakai, Türkay & Yang, 2006
- Species Neohelice granulata (Dana, 1851)
- Genus Paragrapsus H. Milne Edwards, 1853
- Species Paragrapsus gaimardii (H. Milne Edwards, 1837)
- Species Paragrapsus laevis (Dana, 1851)
- Species Paragrapsus quadridentatus (H. Milne Edwards, 1837)
- Species Paragrapsus urvillei H. Milne Edwards, 1853
- Species Paragrapsus verreauxi H. Milne Edwards, 1853 accepted as Paragrapsus laevis (Dana, 1851) (unaccepted > junior subjective synonym)
- Genus Parahelice Sakai, Türkay & Yang, 2006
- Species Parahelice balssi (Sakai, Türkay & Yang, 2006)
- Species Parahelice daviei (Sakai, Türkay & Yang, 2006)
- Species Parahelice georgei (Clark, 1987)
- Species Parahelice ngankeeae Hsu & Shih, 2024
- Species Parahelice pilimana (A. Milne-Edwards, 1873)
- Species Parahelice pilosa (Sakai, Türkay & Yang, 2006)
- Genus Pseudohelice Sakai, Türkay & Yang, 2006
- Species Pseudohelice annamalai Prema, Hsu, Shih & Ravichandran, 2022
- Species Pseudohelice latreillii (H. Milne Edwards, 1837)
- Species Pseudohelice subquadrata (Dana, 1851)
- Subgenus Pseudohelice (Parahelice) Sakai, Türkay & Yang, 2006 accepted as Parahelice Sakai, Türkay & Yang, 2006
- Species Pseudohelice (Parahelice) balssi Sakai, Türkay & Yang, 2006 accepted as Parahelice balssi (Sakai, Türkay & Yang, 2006) (unaccepted > superseded combination)
- Species Pseudohelice (Parahelice) daviei Sakai, Türkay & Yang, 2006 accepted as Parahelice daviei (Sakai, Türkay & Yang, 2006) (unaccepted > superseded combination)
- Species Pseudohelice (Parahelice) pilosa Sakai, Türkay & Yang, 2006 accepted as Parahelice pilosa (Sakai, Türkay & Yang, 2006)
- Genus Gnathochasmus MacLeay, 1838 accepted as Cyclograpsus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Species Gnathochasmus barbatus MacLeay, 1838 accepted as Cyclograpsus barbatus (MacLeay, 1838) (unaccepted > superseded combination)
- Genus Rhaconotus Gerstaecker, 1856 accepted as Metaplax H. Milne Edwards, 1852 (name pre-occupied by Rhaconotus Ruthe, 1854 [Hymenoptera])
- Species Rhaconotus crenulatus Gerstaecker, 1856 accepted as Metaplax crenulata (Gerstaecker, 1856) (unaccepted > superseded combination)
- Subfamily Gaeticinae Davie & Ng, 2007
- Genus Acmaeopleura Stimpson, 1858
- Species Acmaeopleura parvula Stimpson, 1858
- Species Acmaeopleura rotunda Rathbun, 1909
- Species Acmaeopleura balssi Shen, 1932 accepted as Sestrostoma balssi (Shen, 1932) (unaccepted > superseded combination)
- Species Acmaeopleura depressa Sakai, 1965 accepted as Sestrostoma depressum (Sakai, 1965) (unaccepted > superseded combination)
- Species Acmaeopleura hichiro Karasawa & Tanaka, 2005 † accepted as Sestrostoma hichiro (Karasawa & Tanaka, 2005) † (unaccepted > superseded combination)
- Species Acmaeopleura toriumii Takeda, 1974 accepted as Sestrostoma toriumii (Takeda, 1974) (unaccepted > superseded combination)
- Genus Brankocleistostoma Števčić, 2011
- Genus Gaetice Gistel, 1848
- Species Gaetice depressus (De Haan, 1833)
- Species Gaetice ungulatus Sakai, 1939
- Genus Gopkittisak Naruse & Clark, 2009
- Genus Paranotonyx Nobili, 1906
- Genus Proexotelson Naruse, 2015
- Genus Pseudopinnixa Ortmann, 1894
- Genus Sestrostoma Davie & Ng, 2007
- Genus Goetice Gistel, 1848 accepted as Gaetice Gistel, 1848 (ICZN Article 33.2.3.1 used to suppress this original spelling in favor of Gaetice (see Ng et al., 2008) )
- Species Goetice americanus Rathbun, 1923 accepted as Pseudogaetice americanus (Rathbun, 1923) (unaccepted > superseded combination)
- Tribe Gopkittisakini Števčić, 2011 accepted as Gaeticinae Davie & Ng, 2007
- Genus Plalynotus De Haan, 1835 accepted as Platynotus De Haan, 1835 (unaccepted > misspelling - incorrect subsequent spelling)
- Genus Platygrapsus Stimpson, 1858 accepted as Gaetice Gistel, 1848 (unnecessary replacement name for Grapsus (Platynotus) De Haan, 1833)
- Species Platygrapsus convexiusculus Stimpson, 1858 accepted as Gaetice depressus (De Haan, 1833) (unaccepted > junior subjective synonym)
- Species Platygrapsus depressus (De Haan, 1835) accepted as Gaetice depressus (De Haan, 1833) (unaccepted > superseded combination)
- Genus Platynotus De Haan, 1835 accepted as Gaetice Gistel, 1848 (unaccepted > junior homonym)
- Species Platynotus depressus (De Haan, 1833) accepted as Gaetice depressus (De Haan, 1833) (unaccepted > superseded combination)
- Genus Acmaeopleura Stimpson, 1858
- Genus Hemiplax Heller, 1865
- Species Hemiplax hirtipes (Hombron & Jacquinot, 1846)
- Species Hemiplax major Glaessner, 1960 †
- Species Hemiplax boteltobagoe Sakai, 1939 accepted as Macrophthalmus (Paramareotis) boteltobagoe (Sakai, 1939) (unaccepted > superseded combination)
- Species Hemiplax hirtipes (Heller, 1862) accepted as Hemiplax hirtipes (Hombron & Jacquinot, 1846) (unaccepted > junior subjective synonym, also junior secondary homonym)
- Subfamily Pinnotherelinae Alcock, 1900
- Genus Pinnotherelia H. Milne Edwards & Lucas, 1843
- Genus Viapinnixa Schweitzer & Feldmann, 2001 †
- Species Viapinnixa alvarezi Vega, Cosma, Coutiño, Feldmann, Nyborg, Schweitzer & Waugh, 2001 †
- Species Viapinnixa mexiaensis Armstrong, Nyborg, Bishop, Ossó-Morales & Vega, 2009 †
- Species Viapinnixa nodosa (Collins & Rasmussen, 1992) †
- Species Viapinnixa perrilliatae Vega, Nyborg, Fraaye & Espinosa, 2007 †
- Subfamily Thalassograpsinae Davie & Ng, 2007
- Genus Tritodynamia Ortmann, 1894
- Species Tritodynamia bengalensis Trivedi, Mitra & Ng, 2021
- Species Tritodynamia bidentata Yang & Tang, 2005
- Species Tritodynamia dilatata Yang & Sun, 1996
- Species Tritodynamia fujianensis Chen, 1979
- Species Tritodynamia hainanensis Dai in Dai, Feng, Song & Chen, 1980
- Species Tritodynamia horvathi Nobili, 1905
- Species Tritodynamia intermedia Shen, 1935
- Species Tritodynamia japonica Ortmann, 1894
- Species Tritodynamia longipropoda Dai in Dai, Feng, Song & Chen, 1980
- Species Tritodynamia miocaenica Müller, 2006 †
- Species Tritodynamia nontjii Rahayu & Ng, 2021
- Species Tritodynamia rathbuni Shen, 1932
- Species Tritodynamia serratipes Anker & Ng, 2014
- Species Tritodynamia wangi Ng & Ho, 2023
- Species Tritodynamia yeoi Naruse & Ng, 2010
- Species Tritodynamia dilatatum Yang & Sun, 1996 accepted as Tritodynamia dilatata Yang & Sun, 1996 (unaccepted > incorrect grammatical agreement of specific epithet)
- Species Tritodynamia fani Shen, 1932 accepted as Tritodynamia horvathi Nobili, 1905 (unaccepted > junior subjective synonym)
- Species Tritodynamia globosa Karasawa, 1990 † accepted as Asthenognathus globosa (Karasawa, 1990) † (unaccepted > superseded combination)
- Species Tritodynamia longipropodum Dai in Dai, Feng, Song & Chen, 1980 accepted as Tritodynamia longipropoda Dai in Dai, Feng, Song & Chen, 1980 (unaccepted > incorrect grammatical agreement of specific epithet)
- Subfamily Varuninae H. Milne Edwards, 1853
- Genus Brachynotus De Haan, 1833
- Species Brachynotus atlanticus Forest, 1957
- Species Brachynotus corallinus Beschin, Busulini, De Angeli & Tessier, 2007 †
- Species Brachynotus februarius Müller, 1974 †
- Species Brachynotus foresti Zariquiey Álvarez, 1968
- Species Brachynotus gemmellari (Rizza, 1839)
- Species Brachynotus multituberculatus Beschin, Busulini & Tessier in Beschin, Busulini, Tessier & Zorzin, 2019 †
- Species Brachynotus oligocenicus De Angeli, Garassino & Ceccon, 2010 †
- Species Brachynotus sexdentatus (Risso, 1827)
- Subgenus Brachynotus (Heterograpsus) Lucas, 1846 accepted as Brachynotus De Haan, 1833 (unaccepted > junior subjective synonym)
- Species Brachynotus (Heterograpsus) jouyi Rathbun, 1894 accepted as Tetragrapsus jouyi (Rathbun, 1894) (unaccepted > superseded combination)
- Species Brachynotus brevidigitatus Yokoya, 1928 accepted as Hemigrapsus penicillatus (De Haan, 1835) (unaccepted > junior subjective synonym)
- Species Brachynotus edwardsii Hilgendorf, 1882 accepted as Hemigrapsus sexdentatus (H. Milne Edwards, 1837) (unaccepted > junior subjective synonym)
- Species Brachynotus gemmellaroi (Rizza, 1839) accepted as Brachynotus gemmellari (Rizza, 1839) (unaccepted > misspelling - incorrect subsequent spelling)
- Species Brachynotus harpax Hilgendorf, 1892 accepted as Thalassograpsus harpax (Hilgendorf, 1892) (unaccepted > superseded combination)
- Species Brachynotus longitarsis (Miers, 1879) accepted as Hemigrapsus longitarsis (Miers, 1879) (unaccepted > superseded combination)
- Species Brachynotus lucasi (H. Milne Edwards, 1853) accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > junior subjective synonym)
- Species Brachynotus marmoratus (Olivi, 1792) accepted as Pachygrapsus marmoratus (Fabricius, 1787) (unaccepted > superseded combination)
- Species Brachynotus penicillatus (De Haan, 1835) accepted as Hemigrapsus penicillatus (De Haan, 1835) (unaccepted > superseded combination)
- Species Brachynotus spinosus (H. Milne Edwards, 1853) accepted as Parvonotus spinosus (H. Milne Edwards, 1853) (unaccepted > superseded combination)
- Genus Chhapgarus Ng, Trivedi & Bhat, 2022
- Genus Cyrtograpsus Dana, 1851
- Species Cyrtograpsus affinis (Dana, 1851)
- Species Cyrtograpsus altimanus Rathbun, 1914
- Species Cyrtograpsus angulatus Dana, 1851
- Species Cyrtograpsus cirripes Smith, 1869
- Genus Eoacmaeopleura De Angeli & Bellin, 2022 †
- Genus Eriocheir De Haan, 1835
- Species Eriocheir hepuensis Dai, 1991
- Species Eriocheir japonica (De Haan, 1835)
- Species Eriocheir ogasawaraensis Komai in Komai, Yamasaki, Kobayashi, Yamamoto & Watanabe, 2006
- Species Eriocheir sinensis H. Milne Edwards, 1853
- Forma Eriocheir sinensis f. acutifrons Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
- Forma Eriocheir sinensis f. rostrata Panning, 1933 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior objective synonym)
- Forma Eriocheir sinensis f. rotundifrons Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
- Forma Eriocheir sinensis f. trilobata Panning, 1938 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > junior subjective synonym)
- Species Eriocheir formosa Chan, Hung & Yu, 1995 accepted as Platyeriocheir formosa (Chan, Hung & Yu, 1995) (unaccepted > superseded combination)
- Species Eriocheir formosa Nakagawa, 1915 accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > nomen nudum)
- Species Eriocheir japonicus (De Haan, 1835) accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > misspelling - incorrect subsequent spelling)
- Subspecies Eriocheir japonicus hepuensis Dai, 1991 accepted as Eriocheir hepuensis Dai, 1991 (unaccepted > superseded combination)
- Species Eriocheir leptognathus Rathbun, 1913 accepted as Neoeriocheir leptognathus (Rathbun, 1913) (unaccepted > superseded combination)
- Species Eriocheir misakiensis Rathbun, 1919 accepted as Hemigrapsus longitarsis (Miers, 1879) (unaccepted > junior subjective synonym)
- Species Eriocheir rectus Stimpson, 1858 accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > junior subjective synonym)
- Genus Grapsodius Holmes, 1900
- Species Grapsodius eximius Holmes, 1900
- Genus Hemigrapsus Dana, 1851
- Species Hemigrapsus crassimanus Dana, 1851
- Species Hemigrapsus crenulatus (H. Milne Edwards, 1837)
- Species Hemigrapsus estellinensis Creel, 1964
- Species Hemigrapsus gibbus (Hombron & Jacquinot, 1846)
- Species Hemigrapsus longitarsis (Miers, 1879)
- Species Hemigrapsus nudus (Dana, 1851)
- Species Hemigrapsus oregonensis (Dana, 1851)
- Species Hemigrapsus pallipes (H. Milne Edwards, 1837)
- Species Hemigrapsus penicillatus (De Haan, 1835)
- Species Hemigrapsus sanguineus (De Haan, 1835)
- Species Hemigrapsus sexdentatus (H. Milne Edwards, 1837)
- Species Hemigrapsus sinensis Rathbun, 1931
- Species Hemigrapsus takanoi Asakura & Watanabe, 2005
- Species Hemigrapsus affinis Dana, 1851 accepted as Cyrtograpsus affinis (Dana, 1851) (unaccepted > superseded combination)
- Species Hemigrapsus lobulatus Manuel, Gonzales & Basmayor, 1991 accepted as Noarograpsus lobulatus (Manuel, Gonzales & Basmayor, 1991) (unaccepted > superseded combination)
- Species Hemigrapsus tanakoi Asakura & Watanabe, 2005 accepted as Hemigrapsus takanoi Asakura & Watanabe, 2005 (unaccepted > misspelling - incorrect subsequent spelling)
- Genus Neoeriocheir Sakai, 1983
- Genus Noarograpsus Ng, Manuel & Ng, 2006
- Genus Orcovita Ng & Tomascik, 1994
- Species Orcovita angulata Ng, Guinot & Iliffe, 1996
- Species Orcovita fictilia Ng, Guinot & Iliffe, 1996
- Species Orcovita gracilipes Ng, Guinot & Iliffe, 1996
- Species Orcovita hicksi Davie & Ng, 2012
- Species Orcovita holthuisi Ng & Ng, 2009
- Species Orcovita mcneiceae Ng & Ng, 2002
- Species Orcovita miruku Naruse & Tamura, 2006
- Species Orcovita mollitia Ng, Guinot & Iliffe, 1996
- Species Orcovita orchardorum Davie & Ng, 2012
- Species Orcovita saltatrix Ng & Tomascik, 1994
- Species Orcovita tabiacoud Stasolla & Innocenti, 2014
- Species Orcovita varungan Ng & Naruse, 2024
- Genus Otognathon Ng & Števčić, 1993
- Genus Parapyxidognathus Ward, 1941
- Genus Parvonotus Davie & Ng, 2024
- Genus Platyeriocheir Ng, Guo & Ng, 1999
- Species Platyeriocheir formosa (Chan, Hung & Yu, 1995)
- Species Platyeriocheir guangdonga Sakai, 2013 accepted as Platyeriocheir formosa (Chan, Hung & Yu, 1995) (unaccepted > junior subjective synonym)
- Genus Pseudogaetice Davie & Ng, 2007
- Genus Pseudograpsus H. Milne Edwards, 1837
- Species Pseudograpsus albus Stimpson, 1858
- Species Pseudograpsus crassus A. Milne-Edwards, 1868
- Species Pseudograpsus elongatus (A. Milne-Edwards, 1873)
- Species Pseudograpsus setosus (Fabricius, 1798)
- Species Pseudograpsus barbatus H. Milne Edwards, 1853 accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
- Species Pseudograpsus dentatus Miers, 1880 accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
- Species Pseudograpsus erythraeus Kossman, 1877 accepted as Pseudograpsus elongatus (A. Milne-Edwards, 1873) (unaccepted > junior subjective synonym)
- Species Pseudograpsus intermedius Chhapgar, 1957 accepted as Chhapgarus intermedius (Chhapgar, 1957) (unaccepted > superseded combination)
- Species Pseudograpsus laniger Tesch, 1918 accepted as Scutumara laniger (Tesch, 1918) (unaccepted > superseded combination)
- Species Pseudograpsus miyakei Nakamura & Kurata, 1972 accepted as Scutumara miyakei (Nakamura & Takeda, 1972) (unaccepted > superseded combination)
- Species Pseudograpsus nudus Dana, 1851 accepted as Hemigrapsus nudus (Dana, 1851) (unaccepted > superseded combination)
- Species Pseudograpsus oregonensis Dana, 1851 accepted as Hemigrapsus oregonensis (Dana, 1851) (unaccepted > superseded combination)
- Species Pseudograpsus pallipes H. Milne Edwards, 1837 accepted as Hemigrapsus pallipes (H. Milne Edwards, 1837) (unaccepted > superseded combination)
- Species Pseudograpsus penicilliger (Latreille, 1817) accepted as Pseudograpsus setosus (Fabricius, 1798) (unaccepted > junior subjective synonym)
- Genus Ptychognathus Stimpson, 1858
- Species Ptychognathus affinis De Man, 1895
- Species Ptychognathus altimanus (Rathbun, 1914)
- Species Ptychognathus amikee Hsu & Shih, 2024
- Species Ptychognathus andamanensis Pretzmann, 1984
- Species Ptychognathus barbatus (A. Milne-Edwards, 1873)
- Species Ptychognathus capillidigitatus Takeda, 1984
- Species Ptychognathus crassimanus Finnegan, 1931
- Species Ptychognathus demani Roux, 1917
- Species Ptychognathus dentatus De Man, 1892
- Species Ptychognathus easteranus Rathbun, 1907
- Species Ptychognathus glaber Stimpson, 1858
- Subspecies Ptychognathus glaber andamanesis Pretzmann, 1984 accepted as Ptychognathus andamanensis Pretzmann, 1984 (unaccepted > superseded combination)
- Species Ptychognathus guijulugani Rathbun, 1914
- Species Ptychognathus hachijoensis Sakai, 1955
- Species Ptychognathus insolitus Osawa & Ng, 2006
- Species Ptychognathus intermedius (De Man, 1879)
- Species Ptychognathus ishii Sakai, 1939
- Species Ptychognathus johannae Rathbun, 1913
- Species Ptychognathus lipkei Ng, 2010
- Species Ptychognathus makii Hsu & Shih, 2020
- Species Ptychognathus ngankeeae de Mazancourt, Mazel, Marquet, Poupin & Keith, 2024
- Species Ptychognathus onyx Alcock, 1900
- Species Ptychognathus pilipes (A Milne-Edwards, 1868)
- Species Ptychognathus pilosus De Man, 1892
- Species Ptychognathus polleni De Man, 1895
- Species Ptychognathus pusillus Heller, 1865
- Species Ptychognathus riedelii (A Milne-Edwards, 1868)
- Variety Ptychognathus riedelii var. pilosa De Man, 1892 accepted as Ptychognathus pilosus De Man, 1892 (unaccepted > superseded combination)
- Species Ptychognathus sakaii Hsu, Shih & Li, 2022
- Species Ptychognathus similis de Mazancourt, Mazel, Marquet, Poupin & Keith, 2024
- Species Ptychognathus spinicarpus Ortmann, 1894
- Species Ptychognathus stimpsoni Hsu & Shih, 2020
- Species Ptychognathus takahasii Sakai, 1939
- Species Ptychognathus andamanica Alcock, 1900 accepted as Ptychognathus riedelii (A Milne-Edwards, 1868) (unaccepted > junior subjective synonym)
- Species Ptychognathus easterana Rathbun, 1907 accepted as Ptychognathus easteranus Rathbun, 1907 (unaccepted > superseded combination)
- Species Ptychognathus takahashii Sakai, 1939 accepted as Ptychognathus takahasii Sakai, 1939 (unaccepted > misspelling - incorrect subsequent spelling)
- Genus Pyxidognathus A. Milne-Edwards, 1879
- Species Pyxidognathus granulosus A. Milne-Edwards, 1879
- Species Pyxidognathus subglobosus Tesch, 1918
- Species Pyxidognathus deianira De Man, 1888 accepted as Parapyxidognathus deianira (De Man, 1888) (unaccepted > superseded combination)
- Species Pyxidognathus deldeni (De Man, 1879) accepted as Pyxidognathus granulosus A. Milne-Edwards, 1879
- Species Pyxidognathus fluviatilis Alcock, 1900 accepted as Parapyxidognathus fluviatilis (Alcock, 1900) (unaccepted > superseded combination)
- Genus Sakakurapus Karasawa, 2018 †
- Genus Scutumara Ng & Nakasone, 1993
- Genus Tetragrapsus Rathbun, 1918
- Genus Utica White, 1847
- Species Utica barbimana A. Milne-Edwards, 1873
- Species Utica borneensis De Man, 1895
- Species Utica crassimana Haswell, 1881
- Species Utica glabra A. Milne-Edwards, 1873
- Species Utica gracilipes White, 1847
- Species Utica haswelli Wintle, 1887 †
- Species Utica nausithoe De Man, 1895
- Species Utica setosipes Haswell, 1881
- Species Utica yarraensis Wintle, 1887 †
- Species Utica sinensis Parisi, 1918 accepted as Neoeriocheir leptognathus (Rathbun, 1913) (unaccepted > junior subjective synonym)
- Genus Varuna H. Milne Edwards in Bory de Saint Vincent, 1830
- Species Varuna litterata (Fabricius, 1798)
- Species Varuna yui Hwang & Takeda, 1986
- Species Varuna altimana Rathbun, 1914 accepted as Ptychognathus altimanus (Rathbun, 1914) (unaccepted > superseded combination)
- Species Varuna angustifrons Karasawa, 1993 † accepted as Carinocarcinoides angustifrons (Karasawa, 1993) † (unaccepted > superseded combination)
- Species Varuna atlantica Melliss, 1875 accepted as Planes major (MacLeay, 1838) (unaccepted > junior subjective synonym)
- Species Varuna tomentosa Pfeffer, 1889 accepted as Varuna litterata (Fabricius, 1798) (unaccepted > junior subjective synonym)
- Genus Coelochirus Nauck, 1880 accepted as Ptychognathus Stimpson, 1858 (unaccepted > junior subjective synonym)
- Species Coelochirus crinipes Nauck, 1880 accepted as Ptychognathus pilipes (A Milne-Edwards, 1868) (unaccepted > junior subjective synonym)
- Genus Cryptograpsus Dana, 1851 accepted as Cyrtograpsus Dana, 1851 (unaccepted > misspelling - incorrect subsequent spelling)
- Species Cryptograpsus cirripes Smith, 1869 accepted as Cyrtograpsus cirripes Smith, 1869 (unaccepted > superseded combination)
- Genus Eriochirus H. Milne Edwards, 1853 accepted as Eriocheir De Haan, 1835 (unaccepted > unjustified emendation)
- Species Eriochirus japonicus (De Haan, 1835) accepted as Eriocheir japonica (De Haan, 1835) (unaccepted > superseded combination)
- Species Eriochirus sinensis H. Milne Edwards, 1853 accepted as Eriocheir sinensis H. Milne Edwards, 1853 (unaccepted > misspelling - incorrect original spelling)
- Tribe Euchirograpsini Števčić, 2005 accepted as Plagusiidae Dana, 1851
- Genus Gnathograpsus A. Milne-Edwards, 1868 accepted as Ptychognathus Stimpson, 1858 (unaccepted > junior subjective synonym)
- Species Gnathograpsus barbatus A. Milne-Edwards, 1873 accepted as Ptychognathus barbatus (A. Milne-Edwards, 1873) (unaccepted > superseded combination)
- Species Gnathograpsus intermedius De Man, 1879 accepted as Ptychognathus intermedius (De Man, 1879) (unaccepted > superseded combination)
- Species Gnathograpsus pilipes A. Milne-Edwards, 1868 accepted as Ptychognathus pilipes (A Milne-Edwards, 1868) (unaccepted > superseded combination)
- Species Gnathograpsus riedelii A. Milne-Edwards, 1868 accepted as Ptychognathus riedelii (A Milne-Edwards, 1868) (unaccepted > superseded combination)
- Genus Hypsilograpsus De Man, 1879 accepted as Pyxidognathus A. Milne-Edwards, 1879 (unaccepted > junior subjective synonym)
- Species Hypsilograpsus deldeni De Man, 1879 accepted as Pyxidognathus granulosus A. Milne-Edwards, 1879 (unaccepted > junior subjective synonym)
- Tribe Otognathini Števčić, 2011 accepted as Varuninae H. Milne Edwards, 1853
- Genus Pachystomum Nauck, 1880 accepted as Pseudograpsus H. Milne Edwards, 1837 (unaccepted > junior subjective synonym)
- Species Pachystomum philippinense Nauck, 1880 accepted as Pseudograpsus albus Stimpson, 1858 (unaccepted > junior subjective synonym)
- Genus Paraeriocheir Sakai, 2013 accepted as Eriocheir De Haan, 1835 (unaccepted > junior subjective synonym)
- Genus Shurebus Verany, 1846 accepted as Brachynotus De Haan, 1833 (unaccepted > junior subjective synonym)
- Species Shurebus genoensis White, 1847 accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > nomen nudum)
- Species Shurebus genuensis Verany, 1846 accepted as Brachynotus sexdentatus (Risso, 1827) (unaccepted > junior subjective synonym)
- Genus Brachynotus De Haan, 1833
marine, brackish, fresh, terrestrial
Milne Edwards, H. (1853). Mémoire sur la famille des ocypodiens. <em>Annales des Sciences Naturelles, 3rd series.</em> 20: 163-228, pls. 6-11. [details]
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DecaNet eds. (2024). DecaNet. Varunidae H. Milne Edwards, 1853. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=439140 on 2024-07-16
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original description
Milne Edwards, H. (1853). Mémoire sur la famille des ocypodiens. <em>Annales des Sciences Naturelles, 3rd series.</em> 20: 163-228, pls. 6-11. [details]
context source (PeRMS) Moscoso, V. (2012). Catálogo de crustáceos decápodos y estomatópodos del Perú. <em>Boletín Instituto del Mar del Perú.</em> 27(1-2): 8-207., available online at https://repositorio.imarpe.gob.pe/handle/20.500.12958/2190 [details]
![OpenAccess publication OpenAccess publication](https://www.marinespecies.org/aphia/images/openaccess.png)
context source (PeRMS) Moscoso, V. (2012). Catálogo de crustáceos decápodos y estomatópodos del Perú. <em>Boletín Instituto del Mar del Perú.</em> 27(1-2): 8-207., available online at https://repositorio.imarpe.gob.pe/handle/20.500.12958/2190 [details]
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Language | Name | |
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Japanese | モクズガニ科 | [details] |