Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 222, Figs. 8-10: U. Devonian (Frasnian), Russia. 8, Saratov Oblast, sectioned paratype, x 115; 9, 10, Voronezh Oblast, top and side views of holotype, x 100 (from E. V. Bykova, 1952).