Description Loeblich, A. R., Tappan, H. N., 1987: Foraminiferal genera and their classification. Van Nostrand, Reinhold Co. New York 1728 pp. Plate 222, Figs. 1-2: M. Permian (Kungurian), W. Urals, Russia. 1, Diagrammatic section showing inner wall layer (iw) separated from the outer layer (ow) by an intervening space (s) in the apertural region; 2, holotype, x 83 (from Zolotova and Baryshnikov, 1980).