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Ng, N. K., Komai, T., Ng, P. K. L. (2009). Otognathon uru, a new species of crabs (Brachyura, Varunidae) from coral reefs in Okinawa Island, The Ryukyus, Japan. Bulletin of the National Museum of Nature and Science. Series A, Zoology Supplement. 3: 205-214.