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Izuka, T. ; Segawa, S.; Okutani, T. ; Numachi, K. (1994). Evidence on the Existence of Three Species in the Oval Squid Sepioteuthis lessoniana Complex in Ishigaki Island, Okinawa, Southwestern Japan, by Isozyme Analyses. VENUS (The Japanese Journals of Malacology). 53(3):217-228.