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Ahyong, S. T.; Lai, J. C.; Sharkey, D.; Colgan, D. J.; Ng, P. K. L. (2007). Phylogenetics of the brachyuran crabs (Crustacea: Decapoda): The status of Podotremata based on small subunit nuclear ribosomal RNA. Molecular Phylogenetics and Evolution. 45(2): 576–586.