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Schuster, A.; Pisera, A.; Kelly, M.; Bell, L.J.; Pomponi, S.A.; Wörheide, G.; Erpenbeck, D. (2018). New species and a molecular dating analysis of Vetulina Schmidt, 1879 (Porifera: Demospongiae: Sphaerocladina) reveal an ancient relict fauna with Tethys origin. Zoological Journal of the Linnean Society. 184 (3): 585–604.