WoRMS source details
Hayward, B.W.; Holzmann, M.; Pawlowski, J.; Parker, J.H.; Kaushik, T.; Toyofuku, M.S.; Tsuchiya, M. (2021). Molecular and morphological taxonomy of living Ammonia and related taxa (Foraminifera) and their biogeography. Micropaleontology. 67: 109-313.
405477
Hayward, B.W.; Holzmann, M.; Pawlowski, J.; Parker, J.H.; Kaushik, T.; Toyofuku, M.S.; Tsuchiya, M.
2021
Molecular and morphological taxonomy of living Ammonia and related taxa (Foraminifera) and their biogeography
Micropaleontology
67: 109-313.
Publication
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Globally, one of the two most common shallow-marine and estuarine foraminiferal genera is Ammonia. Over the past 50
years, the majority of workers have identified specimens in this genus as belonging to just 1–3 cosmopolitan species – A. beccarii, A.
tepida and A. parkinsoniana. This has been partly because of the problems of discriminating the Ammonia species based entirely on
shell morphology and partly because of a 1974 laboratory study that claimed to have shown that all morphologies were merely
ecophenotypic variants of one species – a conclusion that molecular studies have proven to be unequivocally wrong. In this study we
recognize, describe and figure sixty-seven living species and infraspecies of Recent Ammonia and two closely-related genera
(Acarotrochus, Pseudoeponides) from around the world and summarize their ecological and biogeographic distribution.
Twenty-six species and two subspecies are recognized by DNA sequencing and shown to be morphologically distinguishable. A
further 39 morphospecies and one subspecies, that have not yet been sequenced, have sufficiently distinct morphology to be recognized.
Canonical variates analysis using 42 measured or assessed morphological characters shows that the majority of these species can be
readily discriminated by their test morphologies, although a few of the less-ornamented molecular species are verging on being
pseudocryptic. Molecular sequencing of the type species of Challengerella (C. bradyi) and Helenina (H. anderseni) places them within
our Ammonia clade. Here we continue to recognize the morphologically highly distinct genera Pseudoeponides (subjective senior synonym
of Helenina) and allied Acarotrochus. Twenty new species or subspecies are described (molecular T types in brackets): Ammonia
abramovichae (T8), A. akitaae, A. aoteana australiensis (T5A), A. arabica (T26), A. ariakensis quiltyi, A. buzasi (T11), A. fajemilai, A.
goldsteinae, A. goodayi, A. haigi (T25), A. hattai, A. jorisseni (T23), A. justinparkeri, A. kitazatoi (T10), A. morleyae (T12), A.
shchedrinae, A. turgida almogilabinae (T22M), Acarotrochus lippsi, Pseudoeponides hottingeri and P. dubuissoni. Aneotype is designated
for A. veneta (Schultze 1854) (T1).We recognize 67 Ammonia and related taxa in this study but speculate that there may be 30 or
more additional living species that we are not yet confident to discriminate without molecular sequencing. Ammonia species live in most
parts of the world between 62 °N (Faeroe Islands) and 55 °S (Strait of Magellan), where seasonal sea-surface temperatures are 4–10° C
and above. One estuarine species (A. veneta, T1) is cosmopolitan, euryhaline and eurythermic. Several species are widespread in one or
two ocean regions (e.g., Atlantic and Mediterranean; South Pacific), whereas the majority are endemic to smaller areas (e.g., eastern
Mediterranean; Caribbean-Gulf of Mexico). Eleven biogeographic “provinces” are recognized by cluster analysis of presence/absence
records with the highest diversities in the Australian and northwest Pacific provinces with 18 and 19 species each). Levels of endemism
in our “provinces” range between 0 (temperate Atlantic) and 44% (Australian).
Date
action
by
Acarotrochus Kawagata, 2005 (taxonomy source)
Acarotrochus lippsi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Acarotrochus lobulatus Kawagata, 2005 (taxonomy source)
Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia agoiensis Yanko, 1990 accepted as Ammonia neobeccarii Shchedrina & Mayer, 1975 (source of synonymy)
Ammonia akitaae Hayward, Frenzel & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia angulata (Kuwano, 1950) accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia aoteana subsp. australiensis Hayward & Holzmann, 2021 (original description)
Ammonia arabica Kaushik, 2021 in Hayward et al., 2021 (original description)
Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia asanoi (Uchio, 1951) (taxonomy source)
Ammonia astera Shchedrina, 1984 (taxonomy source)
Ammonia avalonensis (Natland, 1950) (taxonomy source)
Ammonia batava subsp. compacta (Hofker, 1964) accepted as Ammonia batava (Hofker, 1951) (source of synonymy)
Ammonia bulloides (d'Orbigny in Fornasini, 1906) (taxonomy source)
Ammonia buzasi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia catesbyana (d'Orbigny, 1839) (taxonomy source)
Ammonia caucasica Yanko, 1990 accepted as Ammonia karadagica (Dolgopolskaya & Pauli, 1931) (source of synonymy)
Ammonia crebra Shchedrina, 1984 accepted as Ammonia confertitesta Zheng, 1978 (source of synonymy)
Ammonia cristobalensis McCulloch, 1981 (taxonomy source)
Ammonia faceta He, Hu & Wang, 1965 † accepted as Pseudoeponides japonicus Uchio in Kawai et al., 1950 † (additional source)
Ammonia fajemilai Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia flevensis (Hofker, 1930) (taxonomy source)
Ammonia formosa Shchedrina, 1984 accepted as Ammonia confertitesta Zheng, 1978 (source of synonymy)
Ammonia gaudichaudi (d'Orbigny in Fornasini, 1906) (taxonomy source)
Ammonia goldsteinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia goodayi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia granuloumbilica Zheng, 1978 (taxonomy source)
Ammonia haigi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia hattai Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia heterogeneus (Shchedrina, 1984) accepted as Ammonia heterogenea (Shchedrina, 1984) (additional source)
Ammonia jorisseni Hayward, Holzmann & Renjaan, 2021 in Hayward et al., 2021 (original description)
Ammonia juliae McCulloch, 1981 accepted as Ammonia turgida (Hofker, 1951) (source of synonymy)
Ammonia justinparkeri Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia lalibertadensis McCulloch, 1977 accepted as Ammonia decorata Golik, 1977 (source of synonymy)
Ammonia langeri Hayward & Holzmann, 2019 accepted as Pseudoeponides langeri (Hayward & Holzmann, 2019) (additional source)
Ammonia limbatobeccarii (McLean, 1956) † accepted as Ammonia sobrina (Shupack, 1934) (additional source)
Ammonia limnetes (Todd & Brönnimann, 1957) accepted as Ammonia advena (Cushman, 1922) (source of synonymy)
Ammonia morleyae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia multicella Zheng, 1978 (taxonomy source)
Ammonia nantongensis He, Hu & Wang, 1965 (taxonomy source)
Ammonia neobeccarii subsp. huanhajensis Shchedrina & Mayer, 1975 (source of synonymy)
Ammonia novoeuxinica Yanko in Yanko & Troitskaya, 1987 (taxonomy source)
Ammonia pauciloculata subsp. major He, Hu & Wang, 1965 † (taxonomy source)
Ammonia pauperata (Parr, 1941) (taxonomy source)
Ammonia penangensis McCulloch, 1981 (additional source)
Ammonia pseudotepida (Ujiie, 1992) accepted as Ammonia paucipora Zheng, 1979 (source of synonymy)
Ammonia rupta Lei & Li, 2016 accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia takanabensis (Ishizaki, 1943) accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia tepida subsp. bombayensis McCulloch, 1981 (taxonomy source)
Ammonia tepida subsp. cochensis McCulloch, 1981 (taxonomy source)
Ammonia tepida subsp. nitae McCulloch, 1981 accepted as Ammonia aberdoveyensis Haynes, 1973 (source of synonymy)
Ammonia turgida subsp. almogilabinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia umbilicata (d'Orbigny in Fornasini, 1902) (taxonomy source)
Challengerella Billman, Hottinger & Oesterle, 1980 accepted as Ammonia Brünnich, 1771 (source of synonymy)
Cyclohelenina formosaensis Chien, Chang & Yang, 2020 accepted as Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (source of synonymy)
Helenina anderseni (Warren, 1957) accepted as Pseudoeponides anderseni Warren, 1957 (source of synonymy)
Helenina davescottensis van Hengstum & Bernhard, 2016 accepted as Pseudoeponides davescottensis (van Hengstum & Bernhard, 2016) (additional source)
Helenina liujunensis Sun & Zhang in Gou & Tang, 2000 accepted as Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (additional source)
Heleninidae Loeblich & Tappan, 1987 accepted as Ammoniidae Saidova, 1981 (source of synonymy)
Pararotalia capetownensis (McCulloch, 1977) accepted as Pararotalia armata (d'Orbigny in Parker, Jones & Brady, 1865) (source of synonymy)
Pseudoeponides Uchio in Kawai et al., 1950 (additional source)
Pseudoeponides anderseni Warren, 1957 (additional source)
Pseudoeponides angulatus Shchedrina, 1984 accepted as Ammonia veneta (Schultze, 1854) (source of synonymy)
Pseudoeponides compressum Zheng, 1978 accepted as Pseudoeponides anderseni Warren, 1957 (source of synonymy)
Pseudoeponides dubuissoni Baccaert, 2021 in Hayward et al., 2021 (original description)
Pseudoeponides falsobeccarii Rouvillois, 1974 accepted as Ammonia falsobeccarii (Rouvillois, 1974) (source of synonymy)
Pseudoeponides heterogeneus Shchedrina, 1984 accepted as Ammonia heterogeneus (Shchedrina, 1984) accepted as Ammonia heterogenea (Shchedrina, 1984) (additional source)
Pseudoeponides hottingeri Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Pseudoeponides japonicus Uchio in Kawai et al., 1950 † (additional source)
Pseudoeponides ketienziensis (Ishizaki, 1943) accepted as Ammonia ketienziensis (Ishizaki, 1943) (additional source)
Pseudoeponides langeri (Hayward & Holzmann, 2019) (new combination reference)
Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (additional source)
Pseudoeponides whittakeri Bhatia, 1985 accepted as Pseudoeponides anderseni Warren, 1957 (additional source)
Pseudohelenina Collins, 1974 accepted as Ammonia Brünnich, 1771 (additional source)
Pseudohelenina collinsi (Parr, 1932) accepted as Ammonia collinsi (Parr, 1932) (additional source)
Acarotrochus lippsi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Acarotrochus lobulatus Kawagata, 2005 (taxonomy source)
Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia agoiensis Yanko, 1990 accepted as Ammonia neobeccarii Shchedrina & Mayer, 1975 (source of synonymy)
Ammonia akitaae Hayward, Frenzel & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia angulata (Kuwano, 1950) accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia aoteana subsp. australiensis Hayward & Holzmann, 2021 (original description)
Ammonia arabica Kaushik, 2021 in Hayward et al., 2021 (original description)
Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia asanoi (Uchio, 1951) (taxonomy source)
Ammonia astera Shchedrina, 1984 (taxonomy source)
Ammonia avalonensis (Natland, 1950) (taxonomy source)
Ammonia batava subsp. compacta (Hofker, 1964) accepted as Ammonia batava (Hofker, 1951) (source of synonymy)
Ammonia bulloides (d'Orbigny in Fornasini, 1906) (taxonomy source)
Ammonia buzasi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia catesbyana (d'Orbigny, 1839) (taxonomy source)
Ammonia caucasica Yanko, 1990 accepted as Ammonia karadagica (Dolgopolskaya & Pauli, 1931) (source of synonymy)
Ammonia crebra Shchedrina, 1984 accepted as Ammonia confertitesta Zheng, 1978 (source of synonymy)
Ammonia cristobalensis McCulloch, 1981 (taxonomy source)
Ammonia faceta He, Hu & Wang, 1965 † accepted as Pseudoeponides japonicus Uchio in Kawai et al., 1950 † (additional source)
Ammonia fajemilai Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia flevensis (Hofker, 1930) (taxonomy source)
Ammonia formosa Shchedrina, 1984 accepted as Ammonia confertitesta Zheng, 1978 (source of synonymy)
Ammonia gaudichaudi (d'Orbigny in Fornasini, 1906) (taxonomy source)
Ammonia goldsteinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia goodayi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia granuloumbilica Zheng, 1978 (taxonomy source)
Ammonia haigi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia hattai Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia heterogeneus (Shchedrina, 1984) accepted as Ammonia heterogenea (Shchedrina, 1984) (additional source)
Ammonia jorisseni Hayward, Holzmann & Renjaan, 2021 in Hayward et al., 2021 (original description)
Ammonia juliae McCulloch, 1981 accepted as Ammonia turgida (Hofker, 1951) (source of synonymy)
Ammonia justinparkeri Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia lalibertadensis McCulloch, 1977 accepted as Ammonia decorata Golik, 1977 (source of synonymy)
Ammonia langeri Hayward & Holzmann, 2019 accepted as Pseudoeponides langeri (Hayward & Holzmann, 2019) (additional source)
Ammonia limbatobeccarii (McLean, 1956) † accepted as Ammonia sobrina (Shupack, 1934) (additional source)
Ammonia limnetes (Todd & Brönnimann, 1957) accepted as Ammonia advena (Cushman, 1922) (source of synonymy)
Ammonia morleyae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia multicella Zheng, 1978 (taxonomy source)
Ammonia nantongensis He, Hu & Wang, 1965 (taxonomy source)
Ammonia neobeccarii subsp. huanhajensis Shchedrina & Mayer, 1975 (source of synonymy)
Ammonia novoeuxinica Yanko in Yanko & Troitskaya, 1987 (taxonomy source)
Ammonia pauciloculata subsp. major He, Hu & Wang, 1965 † (taxonomy source)
Ammonia pauperata (Parr, 1941) (taxonomy source)
Ammonia penangensis McCulloch, 1981 (additional source)
Ammonia pseudotepida (Ujiie, 1992) accepted as Ammonia paucipora Zheng, 1979 (source of synonymy)
Ammonia rupta Lei & Li, 2016 accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia takanabensis (Ishizaki, 1943) accepted as Ammonia ketienziensis (Ishizaki, 1943) (source of synonymy)
Ammonia tepida subsp. bombayensis McCulloch, 1981 (taxonomy source)
Ammonia tepida subsp. cochensis McCulloch, 1981 (taxonomy source)
Ammonia tepida subsp. nitae McCulloch, 1981 accepted as Ammonia aberdoveyensis Haynes, 1973 (source of synonymy)
Ammonia turgida subsp. almogilabinae Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Ammonia umbilicata (d'Orbigny in Fornasini, 1902) (taxonomy source)
Challengerella Billman, Hottinger & Oesterle, 1980 accepted as Ammonia Brünnich, 1771 (source of synonymy)
Cyclohelenina formosaensis Chien, Chang & Yang, 2020 accepted as Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (source of synonymy)
Helenina anderseni (Warren, 1957) accepted as Pseudoeponides anderseni Warren, 1957 (source of synonymy)
Helenina davescottensis van Hengstum & Bernhard, 2016 accepted as Pseudoeponides davescottensis (van Hengstum & Bernhard, 2016) (additional source)
Helenina liujunensis Sun & Zhang in Gou & Tang, 2000 accepted as Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (additional source)
Heleninidae Loeblich & Tappan, 1987 accepted as Ammoniidae Saidova, 1981 (source of synonymy)
Pararotalia capetownensis (McCulloch, 1977) accepted as Pararotalia armata (d'Orbigny in Parker, Jones & Brady, 1865) (source of synonymy)
Pseudoeponides Uchio in Kawai et al., 1950 (additional source)
Pseudoeponides anderseni Warren, 1957 (additional source)
Pseudoeponides angulatus Shchedrina, 1984 accepted as Ammonia veneta (Schultze, 1854) (source of synonymy)
Pseudoeponides compressum Zheng, 1978 accepted as Pseudoeponides anderseni Warren, 1957 (source of synonymy)
Pseudoeponides dubuissoni Baccaert, 2021 in Hayward et al., 2021 (original description)
Pseudoeponides falsobeccarii Rouvillois, 1974 accepted as Ammonia falsobeccarii (Rouvillois, 1974) (source of synonymy)
Pseudoeponides heterogeneus Shchedrina, 1984 accepted as Ammonia heterogeneus (Shchedrina, 1984) accepted as Ammonia heterogenea (Shchedrina, 1984) (additional source)
Pseudoeponides hottingeri Hayward & Holzmann, 2021 in Hayward et al., 2021 (original description)
Pseudoeponides japonicus Uchio in Kawai et al., 1950 † (additional source)
Pseudoeponides ketienziensis (Ishizaki, 1943) accepted as Ammonia ketienziensis (Ishizaki, 1943) (additional source)
Pseudoeponides langeri (Hayward & Holzmann, 2019) (new combination reference)
Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000) (additional source)
Pseudoeponides whittakeri Bhatia, 1985 accepted as Pseudoeponides anderseni Warren, 1957 (additional source)
Pseudohelenina Collins, 1974 accepted as Ammonia Brünnich, 1771 (additional source)
Pseudohelenina collinsi (Parr, 1932) accepted as Ammonia collinsi (Parr, 1932) (additional source)
Albanian Exclusive Economic Zone for Ammonia beccarii (Linnaeus, 1758)
Albanian Exclusive Economic Zone for Ammonia neobeccarii Shchedrina & Mayer, 1975
Albanian Exclusive Economic Zone for Ammonia parkinsoniana (d'Orbigny, 1839)
Algerian part of the Mediterranean Sea - Western Basin for Ammonia falsobeccarii (Rouvillois, 1974)
Algerian part of the Mediterranean Sea - Western Basin for Ammonia paucipora Zheng, 1979
Algerian part of the Mediterranean Sea - Western Basin for Ammonia veneta (Schultze, 1854)
Angolan part of the South Atlantic Ocean for Ammonia batava (Hofker, 1951)
Anguilla part of the Caribbean Sea for Ammonia batava (Hofker, 1951)
Antigua and Barbuda part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Aral Sea for Ammonia caspica Shchedrina in Mayer, 1968
Argentinean part of the South Atlantic Ocean for Ammonia batava (Hofker, 1951)
Aruban part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Aruban part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Aruban part of the Caribbean Sea for Pseudoeponides anderseni Warren, 1957
Australian part of Bass Strait for Pseudoeponides langeri (Hayward & Holzmann, 2019)
Australian part of the Indian Ocean for Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021
Australian part of the Indian Ocean for Ammonia convexa (Collins, 1958)
Australian part of the Indian Ocean for Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021
Australian part of the Indian Ocean for Ammonia veneta (Schultze, 1854)
Australian part of the Indian Ocean for Pseudoeponides anderseni Warren, 1957
Bahamas part of the North Atlantic Ocean for Pseudoeponides anderseni Warren, 1957
Bahraini part of the Persian Gulf for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Bahraini part of the Persian Gulf for Ammonia veneta (Schultze, 1854)
Belgian part of the North Sea for Ammonia batava (Hofker, 1951)
Bermudian part of the North Atlantic Ocean for Ammonia tepida (Cushman, 1926)
Bermudian part of the North Atlantic Ocean for Pseudoeponides anderseni Warren, 1957
Black Sea for Ammonia batava (Hofker, 1951)
Bonaire part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Bora-Bora for Ammonia ariakensis Akimoto, 2002
Bora-Bora for Ammonia confertitesta Zheng, 1978
Bora-Bora for Ammonia convexa (Collins, 1958)
Brazilian part of the South Atlantic Ocean for Ammonia arabica Kaushik, 2021 in Hayward et al., 2021
Brazilian part of the South Atlantic Ocean for Ammonia pauciloculata (Phleger & Parker, 1951)
Brazilian part of the South Atlantic Ocean for Ammonia rolshauseni (Cushman & Bermúdez, 1946)
Brazilian part of the South Atlantic Ocean for Ammonia tepida (Cushman, 1926)
Brazilian part of the South Atlantic Ocean for Ammonia veneta (Schultze, 1854)
Bulgarian part of the Black Sea for Ammonia beccarii (Linnaeus, 1758)
Bulgarian part of the Black Sea for Ammonia caspica Shchedrina in Mayer, 1968
Bulgarian part of the Black Sea for Ammonia neobeccarii Shchedrina & Mayer, 1975
Bulgarian part of the Black Sea for Ammonia parasovica Shchedrina & Mayer, 1975
Bulgarian part of the Black Sea for Ammonia parkinsoniana (d'Orbigny, 1839)
California for Ammonia veneta (Schultze, 1854)
Canadian part of the North Pacific Ocean for Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021
Caspian Sea for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Caspian Sea for Ammonia caspica Shchedrina in Mayer, 1968
Caspian Sea for Ammonia parkinsoniana (d'Orbigny, 1839)
Chatham Island, New Zealand for Ammonia aoteana (Finlay, 1940)
Chilean part of the South Pacific Ocean for Ammonia aoteana (Finlay, 1940)
Chilean part of the South Pacific Ocean for Ammonia veneta (Schultze, 1854)
Chinese Exclusive Economic Zone for Ammonia confertitesta Zheng, 1978
Chinese Exclusive Economic Zone for Ammonia convexidorsa Zheng, 1978
Chinese Exclusive Economic Zone for Ammonia dilucida Shchedrina, 1984
Chinese Exclusive Economic Zone for Ammonia glans Shchedrina, 1984
Chinese Exclusive Economic Zone for Ammonia heterogeneus (Shchedrina, 1984)
Chinese Exclusive Economic Zone for Ammonia japonica (Hada, 1931)
Chinese Exclusive Economic Zone for Ammonia ketienziensis (Ishizaki, 1943)
Chinese Exclusive Economic Zone for Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021
Chinese Exclusive Economic Zone for Ammonia veneta (Schultze, 1854)
Chinese Exclusive Economic Zone for Pseudoeponides anderseni Warren, 1957
Chinese Exclusive Economic Zone for Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000)
Cook Islands part of the South Pacific Ocean for Acarotrochus lippsi Hayward & Holzmann, 2021 in Hayward et al., 2021
Cook Islands part of the South Pacific Ocean for Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021
Corsica for Ammonia beccarii (Linnaeus, 1758)
Crete for Ammonia beccarii (Linnaeus, 1758)
Croatian Exclusive Economic Zone for Ammonia neobeccarii Shchedrina & Mayer, 1975
Croatian Exclusive Economic Zone for Ammonia paucipora Zheng, 1979
Cuban part of the Caribbean Sea for Ammonia buzasi Hayward & Holzmann, 2021 in Hayward et al., 2021
Cuban part of the Caribbean Sea for Ammonia jacksoni Buzas, Smith & Beem, 1977
Cuban part of the Caribbean Sea for Ammonia veneta (Schultze, 1854)
Curaçaoan part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Curaçaoan part of the Caribbean Sea for Ammonia batava (Hofker, 1951)
Curaçaoan part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Curaçaoan part of the Caribbean Sea for Pseudoeponides anderseni Warren, 1957
Danish part of the North Sea for Ammonia batava (Hofker, 1951)
Danish part of the North Sea for Ammonia confertitesta Zheng, 1978
Danish part of the North Sea for Ammonia veneta (Schultze, 1854)
Dead Sea for Ammonia karadagica (Dolgopolskaya & Pauli, 1931)
Dutch part of the North Sea for Ammonia aberdoveyensis Haynes, 1973
Dutch part of the North Sea for Ammonia batava (Hofker, 1951)
Dutch part of the North Sea for Ammonia confertitesta Zheng, 1978
Dutch part of the North Sea for Ammonia veneta (Schultze, 1854)
Ecuadorean part of the North Pacific Ocean for Ammonia decorata Golik, 1977
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia parkinsoniana (d'Orbigny, 1839)
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia paucipora Zheng, 1979
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia veneta (Schultze, 1854)
Egyptian part of the Red Sea for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Egyptian part of the Red Sea for Challengerella bradyi Billman, Hottinger & Oesterle, 1980
Fijian part of the South Pacific Ocean for Ammonia ariakensis Akimoto, 2002
Fijian part of the South Pacific Ocean for Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021
Fijian part of the South Pacific Ocean for Ammonia convexa (Collins, 1958)
French part of the Mediterranean Sea - Western Basin for Ammonia aberdoveyensis Haynes, 1973
French part of the Mediterranean Sea - Western Basin for Ammonia batava (Hofker, 1951)
French part of the Mediterranean Sea - Western Basin for Ammonia falsobeccarii (Rouvillois, 1974)
French part of the Mediterranean Sea - Western Basin for Ammonia neobeccarii Shchedrina & Mayer, 1975
French part of the Mediterranean Sea - Western Basin for Ammonia venecpeyreae Hayward & Holzmann, 2019
French part of the Mediterranean Sea - Western Basin for Ammonia veneta (Schultze, 1854)
French part of the North Atlantic Ocean for Ammonia aberdoveyensis Haynes, 1973
French part of the North Atlantic Ocean for Ammonia batava (Hofker, 1951)
French part of the North Atlantic Ocean for Ammonia confertitesta Zheng, 1978
Albanian Exclusive Economic Zone for Ammonia neobeccarii Shchedrina & Mayer, 1975
Albanian Exclusive Economic Zone for Ammonia parkinsoniana (d'Orbigny, 1839)
Algerian part of the Mediterranean Sea - Western Basin for Ammonia falsobeccarii (Rouvillois, 1974)
Algerian part of the Mediterranean Sea - Western Basin for Ammonia paucipora Zheng, 1979
Algerian part of the Mediterranean Sea - Western Basin for Ammonia veneta (Schultze, 1854)
Angolan part of the South Atlantic Ocean for Ammonia batava (Hofker, 1951)
Anguilla part of the Caribbean Sea for Ammonia batava (Hofker, 1951)
Antigua and Barbuda part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Aral Sea for Ammonia caspica Shchedrina in Mayer, 1968
Argentinean part of the South Atlantic Ocean for Ammonia batava (Hofker, 1951)
Aruban part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Aruban part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Aruban part of the Caribbean Sea for Pseudoeponides anderseni Warren, 1957
Australian part of Bass Strait for Pseudoeponides langeri (Hayward & Holzmann, 2019)
Australian part of the Indian Ocean for Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021
Australian part of the Indian Ocean for Ammonia convexa (Collins, 1958)
Australian part of the Indian Ocean for Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021
Australian part of the Indian Ocean for Ammonia veneta (Schultze, 1854)
Australian part of the Indian Ocean for Pseudoeponides anderseni Warren, 1957
Bahamas part of the North Atlantic Ocean for Pseudoeponides anderseni Warren, 1957
Bahraini part of the Persian Gulf for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Bahraini part of the Persian Gulf for Ammonia veneta (Schultze, 1854)
Belgian part of the North Sea for Ammonia batava (Hofker, 1951)
Bermudian part of the North Atlantic Ocean for Ammonia tepida (Cushman, 1926)
Bermudian part of the North Atlantic Ocean for Pseudoeponides anderseni Warren, 1957
Black Sea for Ammonia batava (Hofker, 1951)
Bonaire part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Bora-Bora for Ammonia ariakensis Akimoto, 2002
Bora-Bora for Ammonia confertitesta Zheng, 1978
Bora-Bora for Ammonia convexa (Collins, 1958)
Brazilian part of the South Atlantic Ocean for Ammonia arabica Kaushik, 2021 in Hayward et al., 2021
Brazilian part of the South Atlantic Ocean for Ammonia pauciloculata (Phleger & Parker, 1951)
Brazilian part of the South Atlantic Ocean for Ammonia rolshauseni (Cushman & Bermúdez, 1946)
Brazilian part of the South Atlantic Ocean for Ammonia tepida (Cushman, 1926)
Brazilian part of the South Atlantic Ocean for Ammonia veneta (Schultze, 1854)
Bulgarian part of the Black Sea for Ammonia beccarii (Linnaeus, 1758)
Bulgarian part of the Black Sea for Ammonia caspica Shchedrina in Mayer, 1968
Bulgarian part of the Black Sea for Ammonia neobeccarii Shchedrina & Mayer, 1975
Bulgarian part of the Black Sea for Ammonia parasovica Shchedrina & Mayer, 1975
Bulgarian part of the Black Sea for Ammonia parkinsoniana (d'Orbigny, 1839)
California for Ammonia veneta (Schultze, 1854)
Canadian part of the North Pacific Ocean for Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021
Caspian Sea for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Caspian Sea for Ammonia caspica Shchedrina in Mayer, 1968
Caspian Sea for Ammonia parkinsoniana (d'Orbigny, 1839)
Chatham Island, New Zealand for Ammonia aoteana (Finlay, 1940)
Chilean part of the South Pacific Ocean for Ammonia aoteana (Finlay, 1940)
Chilean part of the South Pacific Ocean for Ammonia veneta (Schultze, 1854)
Chinese Exclusive Economic Zone for Ammonia confertitesta Zheng, 1978
Chinese Exclusive Economic Zone for Ammonia convexidorsa Zheng, 1978
Chinese Exclusive Economic Zone for Ammonia dilucida Shchedrina, 1984
Chinese Exclusive Economic Zone for Ammonia glans Shchedrina, 1984
Chinese Exclusive Economic Zone for Ammonia heterogeneus (Shchedrina, 1984)
Chinese Exclusive Economic Zone for Ammonia japonica (Hada, 1931)
Chinese Exclusive Economic Zone for Ammonia ketienziensis (Ishizaki, 1943)
Chinese Exclusive Economic Zone for Ammonia kitazatoi Hayward & Holzmann, 2021 in Hayward et al., 2021
Chinese Exclusive Economic Zone for Ammonia veneta (Schultze, 1854)
Chinese Exclusive Economic Zone for Pseudoeponides anderseni Warren, 1957
Chinese Exclusive Economic Zone for Pseudoeponides liujunensis (Sun & Zhang in Gou & Tang, 2000)
Cook Islands part of the South Pacific Ocean for Acarotrochus lippsi Hayward & Holzmann, 2021 in Hayward et al., 2021
Cook Islands part of the South Pacific Ocean for Ammonia shchedrinae Hayward & Holzmann, 2021 in Hayward et al., 2021
Corsica for Ammonia beccarii (Linnaeus, 1758)
Crete for Ammonia beccarii (Linnaeus, 1758)
Croatian Exclusive Economic Zone for Ammonia neobeccarii Shchedrina & Mayer, 1975
Croatian Exclusive Economic Zone for Ammonia paucipora Zheng, 1979
Cuban part of the Caribbean Sea for Ammonia buzasi Hayward & Holzmann, 2021 in Hayward et al., 2021
Cuban part of the Caribbean Sea for Ammonia jacksoni Buzas, Smith & Beem, 1977
Cuban part of the Caribbean Sea for Ammonia veneta (Schultze, 1854)
Curaçaoan part of the Caribbean Sea for Ammonia advena (Cushman, 1922)
Curaçaoan part of the Caribbean Sea for Ammonia batava (Hofker, 1951)
Curaçaoan part of the Caribbean Sea for Ammonia sobrina (Shupack, 1934)
Curaçaoan part of the Caribbean Sea for Pseudoeponides anderseni Warren, 1957
Danish part of the North Sea for Ammonia batava (Hofker, 1951)
Danish part of the North Sea for Ammonia confertitesta Zheng, 1978
Danish part of the North Sea for Ammonia veneta (Schultze, 1854)
Dead Sea for Ammonia karadagica (Dolgopolskaya & Pauli, 1931)
Dutch part of the North Sea for Ammonia aberdoveyensis Haynes, 1973
Dutch part of the North Sea for Ammonia batava (Hofker, 1951)
Dutch part of the North Sea for Ammonia confertitesta Zheng, 1978
Dutch part of the North Sea for Ammonia veneta (Schultze, 1854)
Ecuadorean part of the North Pacific Ocean for Ammonia decorata Golik, 1977
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia parkinsoniana (d'Orbigny, 1839)
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia paucipora Zheng, 1979
Egyptian part of the Mediterranean Sea - Eastern Basin for Ammonia veneta (Schultze, 1854)
Egyptian part of the Red Sea for Ammonia abramovichae Hayward & Holzmann, 2021 in Hayward et al., 2021
Egyptian part of the Red Sea for Challengerella bradyi Billman, Hottinger & Oesterle, 1980
Fijian part of the South Pacific Ocean for Ammonia ariakensis Akimoto, 2002
Fijian part of the South Pacific Ocean for Ammonia ariakensis subsp. quiltyi Hayward & Holzmann, 2021 in Hayward et al., 2021
Fijian part of the South Pacific Ocean for Ammonia convexa (Collins, 1958)
French part of the Mediterranean Sea - Western Basin for Ammonia aberdoveyensis Haynes, 1973
French part of the Mediterranean Sea - Western Basin for Ammonia batava (Hofker, 1951)
French part of the Mediterranean Sea - Western Basin for Ammonia falsobeccarii (Rouvillois, 1974)
French part of the Mediterranean Sea - Western Basin for Ammonia neobeccarii Shchedrina & Mayer, 1975
French part of the Mediterranean Sea - Western Basin for Ammonia venecpeyreae Hayward & Holzmann, 2019
French part of the Mediterranean Sea - Western Basin for Ammonia veneta (Schultze, 1854)
French part of the North Atlantic Ocean for Ammonia aberdoveyensis Haynes, 1973
French part of the North Atlantic Ocean for Ammonia batava (Hofker, 1951)
French part of the North Atlantic Ocean for Ammonia confertitesta Zheng, 1978
Neotype
Neotype proposed from Lagoon of Venice by Hayward et al. (2012) = T1 phylotype. [details]
Taxonomic remark
Junior synonym in opinion of Hofker (1958) and Schweizer et al. (2011) but discounted by Hayward et al. (2021) [details]
Taxonomic remark
When Pseudoeponides was considered to be a junior synonym of Ammonia, then this species became a secondary homonym ... [details]
Taxonomy
Genetically species of this genus fall within the Ammoniidae clade and the genus has therefore been transferred to ... [details]
